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Monodon

Description

The narwhal (Monodon monoceros), or narwhale, is a medium-sized toothed whale that possesses a large "tusk" from a protruding canine tooth. It lives year-round in the Arctic waters around Greenland, Canada, and Russia. It is one of two living species of whale in the family Monodontidae, along with the beluga whale. The narwhal males are distinguished by a long, straight, helical tusk, which is an elongated upper left canine. The narwhal was one of many species described by Carl Linnaeus in his publication Systema Naturae in 1758.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)2440362https://www.gbif.org/species/2440362
PaleoBioDB ID (PBDB)36775https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=36775
Rankgenus
Taxonomy (GBIF,PBDB)Life : Animalia : Chordata : Mammalia : Cetacea : Monodontidae : Monodon
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-TriploblasticaLankester 1877
-NephrozoaJondelius et al. 2002
-DeuterostomiaGrobben 1908
phylumChordataHaeckel 1847
subphylumVertebrataLamarck 1801
superclassGnathostomataGegenbauer 1874
-Osteichthyes
-SarcopterygiiRomer 1955
subclassDipnotetrapodomorpha
subclassTetrapodomorpha
-Tetrapoda
-ReptiliomorphaSäve-Söderbergh 1934
-AnthracosauriaSäve-Söderbergh 1934
-Batrachosauria
-Cotylosauria
-AmniotaHaeckel 1866
-SynapsidaOsborn 1903
-Therapsida
infraorderCynodontia
-EpicynodontiaHopson and Kitching 2001
infraorderEucynodontiaKemp 1982
-ProbainognathiaHopson 1990
-MammaliamorphaRowe 1988
-MammaliaformesRowe 1988
classMammaliaLinnaeus 1758
orderCetaceaBrisson 1762
-PelagicetiUhen 2008
-NeocetiFordyce and de Muizon 2001
suborderOdontocetiFlower 1867
infraorderDelphinidaMuizon 1984
superfamilyDelphinoideaFlower 1865
familyMonodontidaeGray 1821
genusMonodonLinnæus 1758
Common Namenarwhal
Scientific NameMonodon Linnaeus, 1758
Name Published InLinnæus, Carolus. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Laurentii Salvii, Holmiae. Vol. Tomus I, Editio decima, reformata: i-ii, 1-824.
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Monodongenusbelongs to Cetestated without evidenceLinnaeus, 1758
Monodongenusbelongs to Monodontidaestated without evidenceGray, 1821
Monodongenusbelongs to Phocaeninastated without evidenceGray, 1825
Monodongenusbelongs to Delphinidaestated without evidenceGray, 1846
Monodongenusbelongs to Monoceratinastated without evidenceGray, 1846
Monodongenusbelongs to Delphinidaestated without evidenceGray, 1850
Monodongenusbelongs to Monodontidaestated without evidencePictet, 1853
Monodongenusbelongs to Phocaeninastated without evidenceGray, 1864
Monodongenusbelongs to Delphinidaestated with evidenceGray, 1866
Monodongenusbelongs to Beluginaestated without evidenceFlower, 1867
Monodongenusbelongs to Delphinidaestated with evidenceScammon and Cope, 1869
Monodongenusbelongs to Belugidaestated without evidenceGray, 1870
Monodongenusbelongs to Leucodelphinistated without evidenceBrandt, 1873
Monodongenusbelongs to Delphinidaestated without evidenceLydekker, 1887
Monodongenusbelongs to Delphinapterinaestated without evidenceHay, 1902
Monodongenusbelongs to Delphinidaestated without evidencePalmer, 1904
Monodongenusbelongs to Beluginaestated without evidenceAbel, 1905
Monodongenusbelongs to Delphinidaestated with evidenceTurner, 1912
Monodongenusbelongs to Monodontinaestated without evidenceMiller, 1923
Monodongenusbelongs to Beluginaestated without evidenceZittel, 1925
Monodongenusbelongs to Delphinapteridaestated with evidenceWeber, 1928
Monodongenusbelongs to Delphinidaestated without evidenceKellogg, 1928
Monodongenusbelongs to Monodontinaestated without evidenceHay, 1930
Monodongenusbelongs to Monodontidaestated without evidenceSimpson, 1945
Monodongenusbelongs to Monodontinaestated without evidenceEllerman and Morrisson-Scott, 1951
Monodongenusbelongs to Monodontinaestated with evidenceKasuya, 1973
Monodongenusbelongs to Monodontidaestated without evidenceReeves and Tracey, 1980
Monodongenusbelongs to MonodontidaeimpliedCarroll, 1988
Monodongenusbelongs to Monodontinaestated without evidencede Muizon, 1988
Monodongenusbelongs to Monodontidaestated without evidenceStewart and Stewart, 1989
Monodongenusbelongs to MonodontinaeimpliedMcKenna and Bell, 1997
Monodongenusbelongs to Monodontidaestated without evidenceRice, 1998
Monodongenusbelongs to Monodontinaestated without evidenceFordyce and de Muizon, 2001
Monodongenusbelongs to Cetaceasecond handSepkoski, 2002
Monodongenusbelongs to Monodontidaestated without evidenceGeisler and Sanders, 2003
Monodongenusbelongs to DelphinapteridaeimpliedBuchholtz and Schur, 2004
Monodongenusbelongs to Monodontidaestated without evidenceMead and Brownell, 2005
Monodongenusbelongs to Monodontidaestated with evidenceAgnarsson and May-Collado, 2008
Monodongenusbelongs to Monodontidaestated without evidenceBerta, 2017
Status (PBDB)extant
Taxon Size (PBDB)2
Extant Size (PBDB)2 (100%)
First Recorded Appearance16.0 - 3.6 Ma
Neogene
Environmentmarine,freshwater (based on Odontoceti)
Motilityactively mobile (based on Cetacea)
Dietpiscivore (based on Monodontidae)
Reproductionviviparous (based on Odontoceti)
Ontogenymodification of parts (based on Cetacea)
Taphonomyhydroxylapatite (based on Odontoceti)
Primary Reference (PBDB)C. Linnaeus. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1:1-824
Common Name(s) Narwhals, Narwhal
Wikipediahttps://en.wikipedia.org/wiki/Monodon

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Monodon monoceros
species
accepted (GBIF)
listed (PBDB)
Narhwal132.59 Ma
Pleistocene
0 Ma
Extant

Obsolete Names

NameSourceTaxon RankTaxonomy
Narwalus Lacépède, 1804GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Narwalus
Tachynices Brookes, 1828GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Tachynices
Narhvalus Fischer, 1914GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Narhvalus
Diodon Storr, 1780GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Diodon
Narvalus Rafinesque, 1815GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Narvalus
Monodus Schulze, 1897GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Monodus
Narwhalus Duméril, 1806GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Narwhalus
Narvallus Burnett, 1830GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Narvallus
Ceratodon Brünnich, 1772GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Ceratodon
Ceratodontis Brisson, 1762GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Ceratodontis
Ceratodon Brisson, 1762GBIFgenusAnimalia : Chordata : Mammalia : Cetacea : Monodontidae : Ceratodon

Synonymy List

YearName and Author
1758Monodon Linnaeus p. 75
1762Ceratodon Brisson p. 218
1762Ceratodontis Brisson p. 218
1780Diodon Storr
1804Narwalus Lacépède p. 142
1806Narwhalus Dumeril p. 28
1811Ceratodon Illiger p. 142
1814Narvalus Fischer
1821Monodon Gray p. 310
1825Monodon Gray p. 340
1828Tachynices Brookes p. 44
1846Monodon Gray p. 29
1850Monodon Gray p. 74
1853Monodon Pictet p. 384
1864Monodon Gray p. 246
1866Monodon Gray p. 310
1867Monodon Flower p. 115
1868Monodon Gray p. 9
1869Monodon Scammon and Cope p. 21
1870Monodon Gray p. 773
1873Monodon Brandt p. 231
1887Monodon Lydekker p. 75
1890Monodon Cope p. 609
1891Monodon Newton p. 78
1902Monodon Hay p. 594
1904Monodon Palmer p. 431
1904Narwalus Palmer p. 448
1904Monodon Trouessart p. 772
YearName and Author
1905Monodon Abel p. 129
1912Monodon Turner p. 96
1923Monodon Miller p. 40
1925Monodon Zittel p. 87
1928Monodon Kellogg p. 33 figs. Table 1
1928Monodon Weber p. 389
1930Monodon Hay p. 595
1945Monodon Simpson p. 102
1951Monodon Ellerman and Morrisson-Scott p. 729
1960Monodon Fraser and Purves p. 112 figs. Figure 26
1963Monodon Scheffer and Rice p. 5
1973Monodon Kasuya p. 61
1980Monodon Reeves and Tracey p. 1
1988Monodon Carroll
1988Monodon de Muizon p. 162
1989Monodon Stewart and Stewart p. 1
1997Monodon McKenna and Bell p. 389
1998Monodon Rice p. 97
2001Monodon Fordyce and de Muizon p. 179
2002Monodon Sepkoski, Jr.
2003Monodon Geisler and Sanders p. 28
2004Monodon Buchholtz and Schur p. 384 figs. Figure 1
2005Monodon Mead and Brownell p. 735
2008Monodon Agnarsson and May-Collado p. 981 figs. Fig. 5
2014Monodon Murakami et al. p. 502 figs. Figure 7
2016Monodon Marx et al. p. 136
2017Monodon Berta p. 164

References

Linnaeus C. , Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1, 1-824 doi:10.5962/bhl.title.542
Brisson M. J. , Regnum Animale in classes IX distributum, sive synopsis methodica, Ed. 2
Lacépède B. , Histoire Naturelle des Cetacees
Illiger C. , Prodromus Systematis Mammalium et Avium
Gray J. E. On the natural arrangement of vertebrose animals, The London Medical Repository Monthly Journal and Review 15, 296-310
Gray J. E. An outline of an attempt at the disposition of Mammalia into Tribes and Families, with a list of genera apparently appertaining to each Tribe, Annals of Philosophy, new series 10, 337-344
Gray J. E. On the cetaceous animals, The zoology of the voyage of H. M. S. Erebus and Terror, under the command of Capt. Sir J. C. Ross, R. N., F. R. S., during the years 1839 to 1843 1 & 2, 1-53
Gray J. E. , Catalogue of the Mammalia in the collection of the British Museum, with descriptions of all the species and figures of the genera. Part I. Cetacea, 1-153
Pictet F. -J. , Traité de Paléontologie ou Histoire Naturelle des Animaux Fossiles Considérés dans Leurs Rapports Zoologiques et Géologiques [Treatise of Paleontology or the Natural History of Fossil Animals Considered in their Zoological and Geological Relationships] 1, xiv-584
Gray J. E. On the Cetacea which have been observed in the seas surrounding the British Islands, Proceedings of the Scientific Meetings of the Zoological Society of London 1864, 195-248
Gray J. E. , Catalogue of Seals and Whales in the British Museum, 1-402
Flower W. H. Description of the skeleton of Inia geoffrensis and the skull of Pontoporia blainvillii, with remarks on the systematic position of these animals in the Order Cetacea, Transactions of the Zoological Society of London 6, 87-116
Gray J. E. , Synopsis of the species of whales and dolphins in the Collection of the British Museum, 1-10
Scammon C. M., Cope E. D. On the cetaceans of the western coast of North America, Proceedings of the Academy of Natural Sciences of Philadelphia 22, 13-63
Gray J. E. Notes on the arrangement of the genera of delphinoid whales, Proceedings of the Scientific Meetings of the Zoological Society of London 1870, 772-773
Brandt J. F. Untersuchungen über die fossilen und subfossilen cetaceen Europa's, Mémoires de L'Académie Impériale des Sciences de Saint-Petersbourg, Series 7 20 1, 1-372
Lydekker R. , Catalogue of the Fossil Mammalia in the British Museum (Natural History) 5, 1-345
Newton E. T. The vertebrata of the Pliocene deposits of Britain, Memoirs of the Geological Survey of the United Kingdom, 1-137
Hay O. P. (1902) , Bibliography and Catalogue of the Fossil Vertebrata of North America. Bulletin of the United States Geological Survey 179, 1-868
Palmer T. S. (1904) Index generum mammalium: a list of the genera and families of mammals, North American Fauna 23, 1-984
Trouessart E. L. (1904) , Catalogus Mammalium Tam Viventium Quam Fossilium, Fourth Edition, 1-929
Abel O. (1905) Les Odontocètes du Boldérien (Miocène Supérieur) D'Anvers, Mémoires du Musée royal D'Histoire Naturelle de Belgique 3, 1-155
Miller G. S. (1923) The telescoping of the cetacean skull, Smithsonian Miscellaneous Collections 76 5, 1-70
Hay O. P. (1930) Second Bibliography and Catalogue of the Fossil Vertebrata of North America, Carnegie Institution of Washington 390 II, 1-1074
Simpson G. G. (1945) The principles of classification and a classification of mammals, Bulletin of the American Museum of Natural History 85, 1-350
Ellerman J. R., Morrisson-Scott T. C. S. (1951) , Checklist of Palaearctic and Indian Mammals 1758 to 1946, 1-810
Fraser F. C., Purves P. E. (1960) Hearing in Cetaceans, Bulletin of the British Museum (Natural History) 7 1, 1-140
Scheffer V. B., Rice D. W. (1963) A list of the marine mammals of the world, United States Fish and Wildlife Service Special Scientific Report - Fisheries 431, 1-12
Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698
de Muizon C. (1988) Les relations phylogenetiques des Delphinida (Cetacea, Mammalia), Annales de Paleontologie 74 4, 159-227
McKenna M. C., Bell S. K. (1997) , Classification of Mammals Above the Species Level, 1-640
Rice D. W. (1998) Marine Mammals of the World: Systematics and Distribution, The Society for Marine Mammalogy Special Publication 4, 1-231
Fordyce R. E., de Muizon C. (2001) Evolutionary history of the cetaceans: a review, Secondary Adaptations of Tetrapods to Life in the Water, 169-233
Sepkoski, Jr. J. J. (2002) A compendium of fossil marine animal genera, Bulletins of American Paleontology 363, 1-560
Mead J. G., Brownell R. L. (2005) Order Cetacea, Mammalian Species of the World, 723-743
Agnarsson I., May-Collado L. J. (2008) The phylogeny of Cetartiodactyla: The importance of dense taxon sampling, missing data, and the remarkable promise of cytochrome b to provide reliable species-level phylogenies, Molecular Phylogenetics and Evolution 48, 964-985
Murakami M., Shimada C., et al (2014) Eodelphis kabatensis, a new name for the oldest true dolphin Stenella kabatensis Horikawa, 1977 (Cetacea, Odontoceti, Delphinidae), from the upper Miocene of Japan, and the phylogeny and paleobiogeography of Delphinoidea, Journal of Vertebrate Paleontology 34 3, 491-511 doi:10.1080/02724634.2013.816720
Marx F. G., Lambert O., et al (2016) Cetacean Paleobiology, 1-319
Berta A. (2017) , The Rise of Marine Mammals: 50 Million Years of Evolution, 1-198
Aphia2006 - via The Interim Register of Marine and Nonmarine Genera
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Mead, James G., and Robert L. Brownell, Jr. / Wilson, Don E., and DeeAnn M. Reeder, eds., 2005: Order Cetacea. Mammal Species of the World: A Taxonomic and Geographic Reference, 3rd ed., vol. 1. 723-743. - via Integrated Taxonomic Information System (ITIS)
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Linnaeus, C. (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata [10th revised edition], vol. 1: 824 pp. Laurentius Salvius: Holmiae. - via World Register of Marine Species
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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