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Homolidae

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Description

The family Homolidae, known as carrier crabs or porter crabs, contains 14 genera of marine crabs. They mostly live on the continental slope and continental shelf, and are rarely encountered. Members of the Homolidae have their fifth pereiopods (last pair of walking legs) in a sub-dorsal position, which allows them to hold objects in place over the rear half of the carapace. The objects carried include sponges, black corals and gorgonians, and this behaviour may be a defence mechanism against predators. Some species have been observed carrying living sea urchins in a symbiotic relationship which allows them to benefit from the protection of the urchin's dangerous spikes.

From Wikipedia article at https://en.wikipedia.org/wiki/Homolidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	8045	https://www.gbif.org/species/8045
PaleoBioDB ID (PBDB)	93838	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=93838

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Malacostraca : Decapoda : Homolidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
subphylum	Crustacea	Brünnich 1772
class	Malacostraca	Latreille 1802
superorder	Eucarida	Calman 1904
order	Decapoda	Latreille 1802
-	Meiura	Saint Laurent 1980
infraorder	Brachyura	Latreille 1802
superfamily	Homoloidea	de Haan 1839
family	Homolidae	De Haan 1833

Scientific Name

Homolidae

Name Published In

in Siebold, Fauna japon. (Crustacea) (4): 102

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Homolidae	family	belongs to Brachyura	stated without evidence	de Haan, 1833
Homolidae	family	belongs to Homolidea	stated without evidence	Rathbun, 1926
Homolidae	family	belongs to Homoloidea	stated with evidence	Larghi, 2004
Homolidae	family	belongs to Homoloidea	stated without evidence	Schweitzer et al., 2006

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

30 (32%)

First Recorded Appearance

157 - 152 Ma
Jurassic

Motility

actively mobile (based on Malacostraca)

Vision

well-developed (based on Malacostraca)

Diet

carnivore (based on Decapoda)

Taphonomy

chitin (based on Malacostraca)

Primary Reference (PBDB)

C. E. Schweitzer, R. M. Feldmann, and G. Gonzalez-Barba, V. Cosovic. 2006. New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico. Bulletin of the Mizunami Fossil Museum 33:21-45

Common Name(s)

Carrier Crabs, ホモラ, Häxkrabbor

Wikipedia

https://en.wikipedia.org/wiki/Homolidae

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Antarctidromia ✝ genus	accepted (GBIF) listed (PBDB)		1	23.0 Ma Miocene	23.0 Ma Miocene
Antarctomithrax ✝ genus	accepted (GBIF) listed (PBDB)		1	47.8 Ma Eocene	47.8 Ma Eocene
Dagnaudus genus	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0 Ma Extant
Doerflesia ✝ genus	accepted (GBIF) listed (PBDB)		2	152 Ma Late Jurassic	151 Ma Late Jurassic
Eohomola ✝ genus	accepted (GBIF) listed (PBDB)		1	66.0 Ma Late/Upper Cretaceous	66.0 Ma Late/Upper Cretaceous
Eoprosopon ✝ genus	accepted (GBIF) listed (PBDB)
Gastrodorus ✝ genus	accepted (GBIF) listed (PBDB)		6	157 Ma Jurassic	152 Ma Late Jurassic
Gordonopsis genus	accepted (GBIF)	No associated record in PBDB
Homola genus	accepted (GBIF) listed (PBDB)		4	48.6 Ma Eocene	0 Ma Extant
Homolax genus	accepted (GBIF)	No associated record in PBDB
Homolochunia genus	accepted (GBIF) listed (PBDB)		3	0.01 Ma Pleistocene	0 Ma Extant
Homologenus genus	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0 Ma Extant
Homolomannia genus	accepted (GBIF) listed (PBDB)		2	0.01 Ma Pleistocene	0 Ma Extant
Homolopsis ✝ genus	accepted (GBIF) listed (PBDB)		15	122 Ma Early/Lower Cretaceous	41.3 Ma Eocene
Homolus ✝ genus	accepted (GBIF) listed (PBDB)
Hoplitocarcinus ✝ genus	accepted (GBIF) listed (PBDB)
Ihlopsis genus	accepted (GBIF)	No associated record in PBDB
Lamoha genus	accepted (GBIF) listed (PBDB)		3	0.01 Ma Pleistocene	0 Ma Extant
Latheticocarcinus ✝ genus	accepted (GBIF) listed (PBDB)		5	113 Ma Early/Lower Cretaceous	70.6 Ma Late/Upper Cretaceous
Latreillopsis genus	accepted (GBIF) listed (PBDB)		4	0.01 Ma Pleistocene	0 Ma Extant
Lignihomola ✝ genus	accepted (GBIF) listed (PBDB)
Londinimola ✝ genus	accepted (GBIF) listed (PBDB)
Metahomola ✝ genus	accepted (GBIF) listed (PBDB)		3	93.9 Ma Late/Upper Cretaceous	86.3 Ma Late/Upper Cretaceous
Moloha genus	accepted (GBIF) listed (PBDB)				0 Ma Extant
Notiodromia ✝ genus	accepted (GBIF) listed (PBDB)
Palehomola ✝ genus	accepted (GBIF) listed (PBDB)		1	33.9 Ma Oligocene	33.9 Ma Oligocene
Paromola genus	accepted (GBIF) listed (PBDB)		2	0.01 Ma Pleistocene	0 Ma Extant
Paromolopsis genus	accepted (GBIF) listed (PBDB)		2	0.01 Ma Pleistocene	0 Ma Extant
Preclarocarcinus ✝ genus	accepted (GBIF) listed (PBDB)		1	83.5 Ma Late/Upper Cretaceous	83.5 Ma Late/Upper Cretaceous
Prohomola ✝ genus	accepted (GBIF) listed (PBDB)		3	47.8 Ma Eocene	47.8 Ma Eocene
Rhinodromia ✝ genus	accepted (GBIF) listed (PBDB)		3	113 Ma Early/Lower Cretaceous	113 Ma Early/Lower Cretaceous
Yaldwynopsis genus	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0 Ma Extant
Zygastrocarcinus ✝ genus	accepted (GBIF) listed (PBDB)		2	105 Ma Early/Lower Cretaceous	105 Ma Early/Lower Cretaceous
Paramola bathyalis species	accepted (GBIF)	No associated record in PBDB
Peedeehomola deanbogani species	accepted (GBIF)	No associated record in PBDB

Synonymy List

Year	Name and Author
1833	Homolidae De Haan
1833	Homolidae de Haan
1926	Homolidae Rathbun p. 85
1992	Homolidae Karasawa p. 1248
1993	Homolidae Karasawa p. 40
2004	Homolidae Larghi p. 528
2006	Homolidae Schweitzer et al. p. 24 figs. Table 1
2017	Homolidae Luque et al. p. 15

References

Schweitzer C. E., Feldmann R. M., et al (2006) New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico, Bulletin of the Mizunami Fossil Museum 33, 21-45

Rathbun M. J. (1926) The Fossil Stalk-Eyed Crustacea of the Pacific Slope of North America, United States National Museum Bulletin 138, 1-155

Karasawa H. (1992) Fossil decapod crustaceans from the Manda Group (Middle Eocene), Kyushu, Japan, Transactions and Proceedings of the Paleontological Society of Japan, N.S. 167, 1247-1258

Karasawa H. (1993) Cenozoic Decapod Crustacea from Southwest Japan, Bulletin of the Mizunami Fossil Museum 20, 1-92

Larghi C. (2004) Brachyuran decapod Crustacea from the Upper Cretaceous of Lebanon, Journal of Paleontology 78 3, 528-541

Luque J., Schweitzer C. E., et al (2017) Checklist of fossil decapod crustaceans from tropical America. Part I: Anomura and Brachyura, Nauplius 25, e2017025 doi:10.1590/2358-2936e2017025

De Grave, S., Pentcheff, N.D., Ahyong, S.T., Chan, T.-Y., Crandall, K.A., Dworschak, P.C., Felder, D.L., Feldmann, R.M., Fransen, C.H.J.M., Goulding, L.Y.D., Lemaitre, R., Low, M.E.Y., Martin, J.W., Ng, P.K.L., Schweitzer, C.E., Tan, S.H., Tshudy, D. & Wetzer, R. (2009). A classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology. supplement 21: 1-109. - via The Interim Register of Marine and Nonmarine Genera

Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera

WoRMS (Mar 2013) - via The Interim Register of Marine and Nonmarine Genera

International Commission on Zoological Nomenclature. Opinion 522 Suppression under the Plenary Powers (I) of certain names published by C. S. Rafinesque for genera and species of the orders Decapoda and Stomatopoda (class Crustacea) in the period 1814 ? 1818 and (Ii) of certain specific names currently regarded as senior Subjective synonyms of the names of the type species of Homola Leach, 1815, and Lissa Leach, 1815 respectively both being genera assigned to the first of the foregoing orders.Opinions and declarations rendered by the International Commission on Zoological Nomenclature 19(9), 209-248 (1958) - via Official Lists and Indexes of Names in Zoology

Martin, Joel W., and George E. Davis, 2001: An Updated Classification of the Recent Crustacea. Natural History Museum of Los Angeles, Science Series 39. 124. - via Integrated Taxonomic Information System (ITIS)

Williams, Austin B., Lawrence G. Abele, D. L. Felder, H. H. Hobbs, Jr., R. B. Manning, et al., 1989: Common and Scientific Names of Aquatic Invertebrates from the United States and Canada: Decapod Crustaceans. American Fisheries Society Special Publication 17. 77. - via Integrated Taxonomic Information System (ITIS)

Bowman, Thomas E., and Lawrence G. Abele / Lawrence G. Abele, ed. / Dorothy E. Bliss, ed.-in-chief, 1982: Chapter 1: Classification of the Recent Crustacea. The Biology of Crustacea, vol. 1: Systematics, the Fossil Record, and Biogeography. 1-27. - via Integrated Taxonomic Information System (ITIS)

Türkay, M. (2001). Decapoda, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 284-292 - via World Register of Marine Species

d'Udekem d'Acoz, C. (1999). Inventory and distribution of the decapod crustaceans from the northeastern Atlantic, the Mediterranean and the adjacent continental waters north of 25°N. Collection Patrimoines Naturels, 40. Muséum national d'Histoire naturelle. Paris. ISBN 2-86515-114-10. X, 383 pp. - via World Register of Marine Species

Berggren, Matz (2008) Lista över svenska Malacostraca. Excelfil. - via Dyntaxa. Svensk taxonomisk databas

Martin, J.W., & Davis, G.E. (2001). An updated classification of the recent Crustacea. Science Series, 39. Natural History Museum of Los Angeles County. Los Angeles, CA (USA). 124 pp. - via World Register of Marine Species

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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