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Spring site, Flathead County, Montana, USA

Lat/Long (Decimal)	48.3914,-113.703
Co-ordinates Derivation	stated in text
Given Location	Montana, United States
Mindat.org Region (for given coordinates)	Flathead County, Montana, USA

Collections

Collection	Reference	Stratigraphic Name	Comments	Lithology	Age
Spring site, Middle Fork Flathead River (Kishenehn)	Huber J. T., Greenwalt D. (2011)	Kishenehn - Coal Creek	Middle sequence of the Coal Creek member of the Kishenehn Formation, which has been estimated to be 46.2 +/- 0.4 myo (Lutetian) by 40Ar/39Ar analysis and 43.5 +/- 4.9 myo by fission-track analysis	"shale"	47.8 - 41.3 Ma Eocene

Recorded Fossils

Accepted Name	Hierarchy	Age
Mordellidae family	Animalia : Arthropoda : Insecta : Coleoptera : Mordellidae	47.8 - 41.3 Ma Eocene
Ptenidium kishenehnicum species	Animalia : Arthropoda : Insecta : Coleoptera : Ptiliidae : Ptenidium : *Ptenidium kishenehnicum*	47.8 - 41.3 Ma Eocene
Pinophilini tribe	Animalia : Arthropoda : Insecta : Coleoptera : Staphylinidae : Pinophilini	47.8 - 41.3 Ma Eocene
Tympanophorus greenwalti species	Animalia : Arthropoda : Insecta : Coleoptera : Staphylinidae : Tympanophorus : *Tympanophorus greenwalti*	47.8 - 41.3 Ma Eocene
Bibiodes kishenehnensis species	Animalia : Arthropoda : Insecta : Diptera : Bibionidae : Bibiodes : *Bibiodes kishenehnensis*	47.8 - 41.3 Ma Eocene
Lonchoptera eocenica species	Animalia : Arthropoda : Insecta : Diptera : Lonchopteridae : Lonchoptera : *Lonchoptera eocenica*	47.8 - 41.3 Ma Eocene
Aenigmatias kishenehnensis species	Animalia : Arthropoda : Insecta : Diptera : Phoridae : Aenigmatias : *Aenigmatias kishenehnensis*	47.8 - 41.3 Ma Eocene
Tipula (Trichotipula) fji species	Animalia : Arthropoda : Insecta : Diptera : Tipulidae : Tipula : *Tipula (Trichotipula) fji*	47.8 - 41.3 Ma Eocene
Corixidae family	Animalia : Arthropoda : Insecta : Hemiptera : Corixidae	47.8 - 41.3 Ma Eocene
Camponotites sp. genus	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : *Camponotites*	47.8 - 41.3 Ma Eocene
Crematogaster aurora species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Crematogaster : *Crematogaster aurora*	47.8 - 41.3 Ma Eocene
Dolichoderus dlusskyi species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Dolichoderus : *Dolichoderus dlusskyi*	47.8 - 41.3 Ma Eocene
Eoformica brevipetiola species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Eoformica : *Eoformica brevipetiola*	47.8 - 41.3 Ma Eocene
Eoformica latimedia species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Eoformica : *Eoformica latimedia*	47.8 - 41.3 Ma Eocene
Eoformica pinguis species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Eoformica : *Eoformica pinguis*	47.8 - 41.3 Ma Eocene
Formica annosa species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Formica : *Formica annosa*	47.8 - 41.3 Ma Eocene
Ktunaxia jucunda species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Ktunaxia : *Ktunaxia jucunda*	47.8 - 41.3 Ma Eocene
Lasius glom species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Lasius : *Lasius glom*	47.8 - 41.3 Ma Eocene
Ponerites kishenehne species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Ponerites : *Ponerites kishenehne*	47.8 - 41.3 Ma Eocene
Proiridomyrmex rotundatus species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Proiridomyrmex : *Proiridomyrmex rotundatus*	47.8 - 41.3 Ma Eocene
Protazteca eocenica species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Protazteca : *Protazteca eocenica*	47.8 - 41.3 Ma Eocene
Pseudomyrmex saxulum species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Pseudomyrmex : *Pseudomyrmex saxulum*	47.8 - 41.3 Ma Eocene
Solenopsites abdita species	Animalia : Arthropoda : Insecta : Hymenoptera : Formicidae : Solenopsites : *Solenopsites abdita*	47.8 - 41.3 Ma Eocene
Ichneumonidae family	Animalia : Arthropoda : Insecta : Hymenoptera : Ichneumonidae	47.8 - 41.3 Ma Eocene
Cosmocomoidea greenwalti species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Cosmocomoidea : *Cosmocomoidea greenwalti*	47.8 - 41.3 Ma Eocene
Cosmocomoidea kootenai species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Cosmocomoidea : *Cosmocomoidea kootenai*	47.8 - 41.3 Ma Eocene
Cosmocomoidea rasnitsyni species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Cosmocomoidea : *Cosmocomoidea rasnitsyni*	47.8 - 41.3 Ma Eocene
Eoanaphes stethynioides species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Eoanaphes : *Eoanaphes stethynioides*	47.8 - 41.3 Ma Eocene
Eoeustochus sp. genus	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : *Eoeustochus*	47.8 - 41.3 Ma Eocene
Eoeustochus borchersi species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Eoeustochus : *Eoeustochus borchersi*	47.8 - 41.3 Ma Eocene
Eoeustochus kishenehn species	Animalia : Arthropoda : Insecta : Hymenoptera : Mymaridae : Eoeustochus : *Eoeustochus kishenehn*	47.8 - 41.3 Ma Eocene
Eolestes syntheticus species	Animalia : Arthropoda : Insecta : Odonata : Calopterygidae : Eolestes : *Eolestes syntheticus*	47.8 - 41.3 Ma Eocene

References

Huber J. T., Greenwalt D. (2011) Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera, ZooKeys 130, 473-494

Shockley F. W., Greenwalt D. (2013) Ptenidium kishenehnicum (Coleoptera: Ptiliidae), a New Fossil Described from the Kishenehn Oil Shales, with a Checklist of Previously Known Fossil Ptiliids, Proceedings of the Entomological Society of Washington 115 2, 173-181

Greenwalt D. E., Bechly G. (2014) A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America, Zootaxa 3887, 138-156

LaPolla J. S., Greenwalt D. E. (2015) Fossil ants (Hymenoptera: Formicidae) of the Middle Eocene Kishenehn Formation, Sociobiology 62, 163-174

Greenwalt D., Rose T. R., et al (2016) Preservation of mandibular zinc in a beetle from the Eocene Kishenehn Formation of Montana, USA, Canadian Journal of Earth Sciences 53, 614-621

Brunke A. J., Schillhammer H., et al (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae), Journal of Systematic Palaeontology 15, 1015-1025

Greenwalt D. E., Bickel D. J., et al (2019) Diptera of the middle Eocene Kishenehn Formation. I. Documentation of diversity at the family level, Palaeontologia Electronica 22 2.50A, 1-56

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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