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Barbatia

Description

Barbatia is a genus of "bearded" ark clams, marine bivalve mollusks in the family Arcidae, the ark clams.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)2286193https://www.gbif.org/species/2286193
PaleoBioDB ID (PBDB)16210https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=16210
Rankgenus
Taxonomy (GBIF)Life : Animalia : Mollusca : Bivalvia : Arcida : Arcidae : Barbatia
Taxonomy (PBDB)Life : Animalia : Mollusca : Bivalvia : Arcida : Arcidae : Barbatia
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Spiralia
superphylumLophotrochozoa
phylumMolluscaLinnaeus 1758
classBivalviaLinnaeus 1758
-EubivalviaCarter 2011
subclassAutobranchia
infraclassPteriomorphia
-OstreomorphiFerussac 1822
-ArcioniGray 1854
orderArcida
superfamilyArcoidea
familyArcidaeLamarck 1809
genusBarbatiaGray 1840
Scientific NameBarbatia Gray, 1840
Name Published InGray, John E. 1840.
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Barbatiasubgenusbelongs to Arcastated without evidenceGlenn, 1904
Barbatiagenusbelongs to Arcidaestated without evidenceWoods, 1922
Barbatiasubgenusbelongs to Arcastated without evidenceWeaver and Palmer, 1922
Barbatiasubgenusbelongs to Arcastated with evidenceGrant and Gale, 1931
Barbatiasubgenusbelongs to Arcastated without evidenceLoel and Corey, 1932
Barbatiagenusbelongs to Arcidaestated without evidenceTurner, 1938
Barbatiagenusbelongs to Arcidaestated with evidenceGardner, 1943
Barbatiagenusbelongs to Arcidaestated without evidenceDurham, 1944
Barbatiagenusbelongs to Arcinaestated without evidenceGardner, 1945
Barbatiagenusbelongs to Arcidaestated with evidenceEames, 1951
Barbatiagenusbelongs to Arcinaestated with evidenceVokes, 1980
Barbatiagenusbelongs to Arcinaestated without evidenceDockery, 1982
Barbatiagenusbelongs to Arcidaestated without evidenceSquires, 1988
Barbatiagenusbelongs to Arcinaestated with evidencePacaud and Le Renard, 1995
Barbatiagenusbelongs to ArcidaeimpliedLamprell and Healy, 1998
Barbatiagenusbelongs to Arcinaestated with evidenceCoan et al., 2000
Barbatiagenusbelongs to Arcidaestated with evidenceSquires, 2001
Barbatiagenusbelongs to Arcidaestated without evidenceTodd, 2001
Barbatiagenusbelongs to Arcinaestated without evidenceTiwari, 2001
Barbatiagenusbelongs to Arcoidasecond handSepkoski, 2002
Barbatiagenusbelongs to Arcinaestated without evidenceSpencer et al., 2004
Barbatiagenusbelongs to Arcidaestated without evidenceEl Qot, 2006
Barbatiagenusbelongs to Arcinaestated without evidenceMatsubara et al., 2010
Barbatiagenusbelongs to Barbatiidaestated without evidenceBouchet et al., 2010
Status (PBDB)extant
Taxon Size (PBDB)103
Extant Size (PBDB)23 (22%)
First Recorded Appearance242 - 235 Ma
Triassic
Environmentlagoonal,coastal,inner shelf,outer shelf,oceanic,brackish,freshwater (based on Arcidae)
Motilitystationary, attached
Visionlimited (based on Arcidae)
Dietsuspension feeder
Ontogenyaccretion (based on Arcidae)
Ecospace CommentsLIFE HISTORY: EP, SE, BA, SU according to Todd (2001).
Taphonomyaragonite,intermediate,minor folds, major ribbing (based on Arcidae)
Primary Reference (PBDB)J. J. Sepkoski, Jr. 2002. A compendium of fossil marine animal genera. Bulletins of American Paleontology 363:1-560
Common Name(s) エガイ
Wikipediahttps://en.wikipedia.org/wiki/Barbatia

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Barbatia (Barbatia)
subgenus
listed (PBDB)194161 Ma
Jurassic
0 Ma
Extant
Barbatia (Cucullaearca)
subgenus
listed (PBDB)11372.1 Ma
Late/Upper Cretaceous
0 Ma
Extant
Barbatia (Diluvarca)
subgenus
listed (PBDB)
Barbatia (Granoarca)
subgenus
listed (PBDB)0 Ma
Extant
Barbatia (Hawaiarca)
subgenus
listed (PBDB)1320.4 Ma
Miocene
0 Ma
Extant
Barbatia (Mesocibota)
subgenus
listed (PBDB)65.33 Ma
Miocene
0 Ma
Extant
Barbatia (Obliquarca)
subgenus
listed (PBDB)1016.0 Ma
Miocene
0 Ma
Extant
Barbatia (Rostarca)
subgenus
listed (PBDB)466.0 Ma
Late/Upper Cretaceous
41.3 Ma
Eocene
Arca (Barbatia) cowlitzensis
species
listed (PBDB)141.3 Ma
Eocene
41.3 Ma
Eocene
Arca (Barbatia) gracilicostata
species
listed (PBDB)
Arca (Barbatia) landesi
species
listed (PBDB)566.0 Ma
Late/Upper Cretaceous
41.3 Ma
Eocene
Arca (Barbatia) nanggulanensis
species
listed (PBDB)
Arca (Barbatia) oostinghi
species
listed (PBDB)
Arca (Barbatia) strongi
species
listed (PBDB)423.0 Ma
Miocene
23.0 Ma
Miocene
Arca (Barbatia) suzzalloi
species
listed (PBDB)241.3 Ma
Eocene
41.3 Ma
Eocene
Arca (Barbatia) tenuicrenata
species
listed (PBDB)
Arca aegyptiaca
species
accepted (GBIF)No associated record in PBDB
Arca appendiculata
species
accepted (GBIF)No associated record in PBDB
Arca axinaea
species
accepted (GBIF)No associated record in PBDB
Arca carolinensis
species
accepted (GBIF)
listed (PBDB)
1283.6 Ma
Late/Upper Cretaceous
3.60 Ma
Pliocene
Arca cowlitzensis
species
accepted (GBIF)No associated record in PBDB
Arca gracilicostata
species
accepted (GBIF)No associated record in PBDB
Arca landesi
species
accepted (GBIF)No associated record in PBDB
Arca lignitifera
species
accepted (GBIF)No associated record in PBDB
Arca malaiana
species
accepted (GBIF)No associated record in PBDB
Arca mauryae
species
accepted (GBIF)No associated record in PBDB
Arca nanggulanensis
species
accepted (GBIF)No associated record in PBDB
Arca oostinghi
species
accepted (GBIF)No associated record in PBDB
Arca propatula
species
accepted (GBIF)No associated record in PBDB
Arca rembangensis
species
accepted (GBIF)No associated record in PBDB
Arca strongi
species
accepted (GBIF)No associated record in PBDB
Arca subtrigonalis
species
accepted (GBIF)No associated record in PBDB
Arca sundaiana
species
accepted (GBIF)No associated record in PBDB
Arca suzzalloi
species
accepted (GBIF)No associated record in PBDB
Arca tenuicrenata
species
accepted (GBIF)No associated record in PBDB
Barbatia aegyptiaca
species
listed (PBDB)6101 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Barbatia agnastha
species
accepted (GBIF)No associated record in PBDB
Barbatia aldrichi
species
accepted (GBIF)No associated record in PBDB
Barbatia amygdalumtortum
species
accepted (GBIF)No associated record in PBDB
Barbatia amygdalumtostum
species
accepted (GBIF)No associated record in PBDB
Barbatia aspera
species
accepted (GBIF)
listed (PBDB)
347.8 Ma
Eocene
47.8 Ma
Eocene
Barbatia assawomanensis
species
accepted (GBIF)No associated record in PBDB
Barbatia awamoana
species
accepted (GBIF)No associated record in PBDB
Barbatia axinaea
species
listed (PBDB)111.6 Ma
Miocene
11.6 Ma
Miocene
Barbatia barbata
species
accepted (GBIF)No associated record in PBDB
Barbatia barbulata
species
accepted (GBIF)No associated record in PBDB
Barbatia biali
species
accepted (GBIF)No associated record in PBDB
Barbatia bicolorata
species
accepted (GBIF)No associated record in PBDB
Barbatia bonaczyi
species
accepted (GBIF)No associated record in PBDB
Barbatia botanica
species
accepted (GBIF)No associated record in PBDB
Barbatia bullata
species
accepted (GBIF)No associated record in PBDB
Barbatia cahua
species
accepted (GBIF)No associated record in PBDB
Barbatia candida
species
accepted (GBIF)No associated record in PBDB
Barbatia cauderanensis
species
accepted (GBIF)No associated record in PBDB
Barbatia cliffensis
species
accepted (GBIF)
listed (PBDB)
455.8 Ma
Eocene
47.8 Ma
Eocene
Barbatia coalvillensis
species
accepted (GBIF)No associated record in PBDB
Barbatia coelata
species
accepted (GBIF)No associated record in PBDB
Barbatia coma
species
accepted (GBIF)No associated record in PBDB
Barbatia cometa
species
accepted (GBIF)No associated record in PBDB
Barbatia complanata
species
accepted (GBIF)No associated record in PBDB
Barbatia consutilis
species
accepted (GBIF)
listed (PBDB)
Barbatia corvamnis
species
accepted (GBIF)
listed (PBDB)
241.3 Ma
Eocene
38.0 Ma
Eocene
Barbatia costellata
species
accepted (GBIF)No associated record in PBDB
Barbatia crustata
species
accepted (GBIF)
listed (PBDB)
Barbatia culleni
species
accepted (GBIF)
listed (PBDB)
123.0 Ma
Miocene
23.0 Ma
Miocene
Barbatia cuniculana
species
accepted (GBIF)No associated record in PBDB
Barbatia dasia
species
accepted (GBIF)No associated record in PBDB
Barbatia delepida
species
accepted (GBIF)No associated record in PBDB
Barbatia deusseni
species
accepted (GBIF)
listed (PBDB)
141.3 Ma
Eocene
41.3 Ma
Eocene
Barbatia disclusa
species
accepted (GBIF)No associated record in PBDB
Barbatia dissimilis
species
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Barbatia divaricata
species
accepted (GBIF)No associated record in PBDB
Barbatia domingensis
species
accepted (GBIF)No associated record in PBDB
Barbatia dubia
species
accepted (GBIF)No associated record in PBDB
Barbatia endomena
species
accepted (GBIF)No associated record in PBDB
Barbatia foliata
species
accepted (GBIF)
listed (PBDB)
35.33 Ma
Miocene
0 Ma
Extant
Barbatia gabonensis
species
accepted (GBIF)No associated record in PBDB
Barbatia gibba
species
accepted (GBIF)
listed (PBDB)
311.6 Ma
Miocene
2.59 Ma
Pleistocene
Barbatia gravesii
species
accepted (GBIF)No associated record in PBDB
Barbatia grayana
species
accepted (GBIF)No associated record in PBDB
Barbatia hachijoensis
species
accepted (GBIF)No associated record in PBDB
Barbatia halidonta
species
accepted (GBIF)No associated record in PBDB
Barbatia hawaia
species
accepted (GBIF)No associated record in PBDB
Barbatia helblingi
species
listed (PBDB)20.13 Ma
Pleistocene
0.13 Ma
Pleistocene
Barbatia hiloa
species
accepted (GBIF)No associated record in PBDB
Barbatia ickeae
species
accepted (GBIF)No associated record in PBDB
Barbatia illota
species
listed (PBDB)22.59 Ma
Pleistocene
0 Ma
Extant
Barbatia inglisia
species
accepted (GBIF)
listed (PBDB)
338.0 Ma
Eocene
38.0 Ma
Eocene
Barbatia inusitata
species
accepted (GBIF)No associated record in PBDB
Barbatia irregularis
species
doubtful (GBIF)
listed (PBDB)
247.8 Ma
Eocene
47.8 Ma
Eocene
Barbatia islopa
species
accepted (GBIF)No associated record in PBDB
Barbatia jacksonensis
species
accepted (GBIF)No associated record in PBDB
Barbatia kayalensis
species
accepted (GBIF)
listed (PBDB)
1323.0 Ma
Miocene
23.0 Ma
Miocene
Barbatia kickapooensis
species
accepted (GBIF)No associated record in PBDB
Barbatia lacerata
species
accepted (GBIF)No associated record in PBDB
Barbatia laterosa
species
accepted (GBIF)No associated record in PBDB
Barbatia legayi
species
accepted (GBIF)
listed (PBDB)
2168 Ma
Middle Jurassic
168 Ma
Middle Jurassic
Barbatia legumen
species
accepted (GBIF)No associated record in PBDB
Barbatia lesbaritzensis
species
accepted (GBIF)No associated record in PBDB
Barbatia leupoldi
species
accepted (GBIF)No associated record in PBDB
Barbatia lignitifera
species
listed (PBDB)156.0 Ma
Paleocene
56.0 Ma
Paleocene
Barbatia limatella
species
accepted (GBIF)
listed (PBDB)
548.6 Ma
Eocene
37.2 Ma
Eocene
Barbatia ludoviciana
species
accepted (GBIF)
listed (PBDB)
2761.6 Ma
Paleocene
38.0 Ma
Eocene
Barbatia lurida
species
accepted (GBIF)No associated record in PBDB
Barbatia malaiana
species
listed (PBDB)116.0 Ma
Miocene
16.0 Ma
Miocene
Barbatia marylandica
species
listed (PBDB)1933.9 Ma
Oligocene
13.8 Ma
Miocene
Barbatia mauryae
species
listed (PBDB)35.33 Ma
Miocene
3.60 Ma
Pliocene
Barbatia melaviensis
species
accepted (GBIF)No associated record in PBDB
Barbatia mella
species
accepted (GBIF)No associated record in PBDB
Barbatia meridiana
species
accepted (GBIF)
listed (PBDB)
183.6 Ma
Late/Upper Cretaceous
83.6 Ma
Late/Upper Cretaceous
Barbatia merriami
species
accepted (GBIF)No associated record in PBDB
Barbatia microtera
species
accepted (GBIF)No associated record in PBDB
Barbatia molokaia
species
accepted (GBIF)No associated record in PBDB
Barbatia morsei
species
accepted (GBIF)
listed (PBDB)
447.8 Ma
Eocene
47.8 Ma
Eocene
Barbatia multivillosa
species
accepted (GBIF)No associated record in PBDB
Barbatia mundeformata
species
accepted (GBIF)
listed (PBDB)
121.7 Ma
Miocene
21.7 Ma
Miocene
Barbatia nattheimensis
species
accepted (GBIF)No associated record in PBDB
Barbatia nigeriensis
species
accepted (GBIF)No associated record in PBDB
Barbatia nodai
species
listed (PBDB)247.8 Ma
Eocene
47.8 Ma
Eocene
Barbatia novaezealandiae
species
accepted (GBIF)No associated record in PBDB
Barbatia oahua
species
accepted (GBIF)No associated record in PBDB
Barbatia obliquata
species
accepted (GBIF)No associated record in PBDB
Barbatia opthanta
species
accepted (GBIF)No associated record in PBDB
Barbatia paradiagona
species
accepted (GBIF)
listed (PBDB)
333.9 Ma
Oligocene
33.9 Ma
Oligocene
Barbatia parallela
species
accepted (GBIF)No associated record in PBDB
Barbatia parva
species
accepted (GBIF)No associated record in PBDB
Barbatia parvivillosa
species
accepted (GBIF)No associated record in PBDB
Barbatia perinesa
species
accepted (GBIF)No associated record in PBDB
Barbatia perplura
species
accepted (GBIF)No associated record in PBDB
Barbatia phalacra
species
accepted (GBIF)No associated record in PBDB
Barbatia picota
species
accepted (GBIF)No associated record in PBDB
Barbatia pistachia
species
accepted (GBIF)
listed (PBDB)
20.13 Ma
Pleistocene
0.13 Ma
Pleistocene
Barbatia platei
species
accepted (GBIF)No associated record in PBDB
Barbatia propertua
species
accepted (GBIF)No associated record in PBDB
Barbatia pseudoillota
species
listed (PBDB)33.60 Ma
Pliocene
0 Ma
Extant
Barbatia pseudonavicula
species
accepted (GBIF)No associated record in PBDB
Barbatia pumila
species
accepted (GBIF)
listed (PBDB)
116.0 Ma
Miocene
16.0 Ma
Miocene
Barbatia pyrrhotus
species
accepted (GBIF)No associated record in PBDB
Barbatia quilonensis
species
accepted (GBIF)
listed (PBDB)
123.0 Ma
Miocene
23.0 Ma
Miocene
Barbatia reeveana
species
accepted (GBIF)No associated record in PBDB
Barbatia reinharti
species
accepted (GBIF)No associated record in PBDB
Barbatia rembangensis
species
listed (PBDB)116.0 Ma
Miocene
16.0 Ma
Miocene
Barbatia revelata
species
accepted (GBIF)No associated record in PBDB
Barbatia rhombicula
species
accepted (GBIF)No associated record in PBDB
Barbatia riculata
species
accepted (GBIF)No associated record in PBDB
Barbatia sacyi
species
accepted (GBIF)No associated record in PBDB
Barbatia scazon
species
accepted (GBIF)No associated record in PBDB
Barbatia sculpta
species
accepted (GBIF)No associated record in PBDB
Barbatia sculpturata
species
accepted (GBIF)No associated record in PBDB
Barbatia seraperta
species
accepted (GBIF)
listed (PBDB)
247.8 Ma
Eocene
41.3 Ma
Eocene
Barbatia setigera
species
accepted (GBIF)No associated record in PBDB
Barbatia simondsi
species
accepted (GBIF)No associated record in PBDB
Barbatia simulans
species
accepted (GBIF)
listed (PBDB)
Barbatia sociella
species
accepted (GBIF)No associated record in PBDB
Barbatia solidula
species
accepted (GBIF)No associated record in PBDB
Barbatia sorongensis
species
accepted (GBIF)No associated record in PBDB
Barbatia stearnsii
species
accepted (GBIF)No associated record in PBDB
Barbatia subtrigonalis
species
listed (PBDB)111.6 Ma
Miocene
11.6 Ma
Miocene
Barbatia sundaiana
species
listed (PBDB)116.0 Ma
Miocene
16.0 Ma
Miocene
Barbatia supracretacea
species
accepted (GBIF)No associated record in PBDB
Barbatia terebrans
species
accepted (GBIF)No associated record in PBDB
Barbatia terraquetra
species
accepted (GBIF)No associated record in PBDB
Barbatia thomasensis
species
accepted (GBIF)No associated record in PBDB
Barbatia trapezina
species
accepted (GBIF)No associated record in PBDB
Barbatia uhligi
species
accepted (GBIF)No associated record in PBDB
Barbatia uxorispalmeri
species
accepted (GBIF)
listed (PBDB)
1047.8 Ma
Eocene
41.3 Ma
Eocene
Barbatia virescens
species
accepted (GBIF)No associated record in PBDB
Barbatia wendti
species
listed (PBDB)116.0 Ma
Miocene
16.0 Ma
Miocene
Barbatia woodfordi
species
accepted (GBIF)No associated record in PBDB
Barbatia wordeni
species
accepted (GBIF)No associated record in PBDB
Byssoarca marylandica
species
accepted (GBIF)No associated record in PBDB
Scapharca halidonata
species
accepted (GBIF)No associated record in PBDB

Obsolete Names

NameSourceTaxon RankTaxonomy
Savignyarca Jousseaume, 1891GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Savignyarca
Cucullaearca Conrad, 1865GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Cucullaearca
Granoarca Conrad, 1862GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Granoarca
Barbotia Dalton, 1880GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Barbotia
Ustularca Iredale, 1939GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Ustularca
Barbatirus Iredale, 1939GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Barbatirus
Abarbatia Dall, Bartsch & Rehder, 1938GBIFgenusAnimalia : Mollusca : Bivalvia : Arcida : Arcidae : Abarbatia

Synonymy List

YearName and Author
1842Barbatia Gray
1904Arca (Barbatia) Glenn p. 391
1922Arca (Barbatia) Weaver and Palmer p. 8
1922Barbatia Woods p. 62
1931Arca (Barbatia) Grant and Gale p. 143
1932Arca (Barbatia) Loel and Corey
1935Arca (Barbatia) Oostingh p. 125
1938Abarbatia Dall et al.
1938Barbatia Turner p. 42
1939Barbatia Mukerjee p. 25
1943Barbatia Gardner p. 22
1944Barbatia Durham p. 136
1945Barbatia Gardner p. 52
1951Barbatia Eames p. 329
1972Barbatia Hertlein and Grant p. 157
1980Barbatia Vokes p. 11
YearName and Author
1982Barbatia Dockery p. 30
1982Barbatia Yamani
1987Barbatia Squires p. 55
1988Barbatia Squires p. 17
1992Barbatia Squires and Demetrion p. 36
1995Barbatia Pacaud and Le Renard p. 175
1998Barbatia Lamprell and Healy p. 44
2000Barbatia Coan et al. p. 128
2001Barbatia Squires p. 15
2001Barbatia Tiwari p. 150
2001Barbatia Todd
2002Barbatia Sepkoski, Jr.
2004Barbatia Spencer et al.
2006Barbatia El Qot p. 22
2010Barbatia Bouchet et al.
2010Barbatia Matsubara et al. p. 132

References

Sepkoski, Jr. J. J. (2002) A compendium of fossil marine animal genera, Bulletins of American Paleontology 363, 1-560
Glenn L. C. (1904) Pelecypoda, Maryland Geological Survey Miocene Text, 274-401
Weaver C. E., Palmer K. (1922) Fauna from the Eocene of Washington, University of Washington Publications in Geology 1 3, 1-56
Woods H. (1922) Section A. Mollusca from the Eocene and Miocene deposits of Peru. , Geology of the Tertiary and Quaternary Periods in the Northwest Part of Peru, 51-113
Grant U. S., Gale H. R. (1931) Catalogue of the Marine Pliocene and Pleistocene Mollusca of California, Memoirs of the San Diego Society of Natural History 1, 1-1036
Loel W., Corey W. H. (1932) The Vaqueros Formation, Lower Miocene of California (Part 1), University of California Publications, Bulletin of the Department of Geological Sciences 22 3, 31-410
Oostingh C. H. (1935) Die Mollusken des Pliozäns von Boemiajoe (Java), Dienst van den Mijnbouw in Nederlandsch-Indië Wetenschappelijke Mededeelingen 26, 1-247
Turner F. E. (1938) Stratigraphy and Mollusca of the Eocene of western Oregon, Geological Society of America Special Paper 10, 1-130
Mukerjee P. N. (1939) Fossil fauna from the Tertiary of Garo Hills, Assam, Palaeontologia Indica 28 1, 1-101
Gardner J. A. (1943) Mollusca from the Miocene and Lower Pliocene of Virginia and North Carolina, United States Geological Survey Professional Paper 199 A
Durham J. W. (1944) Megafaunal zones of the Oligocene of northwestern Washington, University of California Publications in Geological Sciences 27, 101 -212
Gardner J. (1945) Mollusca of the Tertiary formations of Northeastern Mexico, Geological Society of American Memoir 11
Eames F. E. (1951) A Contribution to the study of the Eocene in Western Pakistan and Western India: B. Description of the Lamellibranchia from standard sections in the Rakni Nala and Zinda Pir areas of the Western Punjab and in the Kohat District, Philosophical Transactions of the Royal Society of London Series B 235, 311-482
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Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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