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Calymenidae ✝

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Calymene celebra

Milwaukee County, Wisconsin, USA

Calymene celebra

Milwaukee County, Wisconsin, USA

Calymene celebra

Milwaukee County, Wisconsin, USA

Calymene celebra

Milwaukee County, Wisconsin, USA

Calymene celebra

Milwaukee County, Wisconsin, USA

Calymene celebra

Milwaukee County, Wisconsin, USA

Description

Calymenidae is a family of trilobites, containing the following genera:

From Wikipedia article at https://en.wikipedia.org/wiki/Calymenidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3266762	https://www.gbif.org/species/3266762
PaleoBioDB ID (PBDB)	85380	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=85380

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Trilobita : Phacopida : Calymenidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
class	Trilobita	Walch 1771
order	Phacopida	Salter 1864
suborder	Calymenina	Swinnerton 1915
family	Calymenidae	Milne-Edwards 1840

Scientific Name

Calymenidae

Name Published In

Histoire naturelle des Crustacés, in: Roret’s Suites à Buffon, vol. 3, p. 293

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Calymenidae	family	belongs to Calymenina	stated without evidence	Harrington and Leanza, 1957
Calymenidae	family	belongs to Agnostida	stated without evidence	Robison and Pantoja-Alor, 1968
Calymenidae	family	belongs to Phacopida	stated without evidence	Dean, 1983
Calymenidae	family	belongs to Phacopida	stated with evidence	Whittington et al., 1997
Calymenidae	family	belongs to Calymenina	stated with evidence	Adrain, 2011

Status (PBDB)

extinct

Taxon Size (PBDB)

137

First Recorded Appearance

485 - 478 Ma
Paleozoic

Last Recorded Appearance

408 - 383 Ma
Devonian

Environment

marine (based on Trilobita)

Motility

actively mobile

Vision

well-developed

Diet

carnivore

Ontogeny

molting,addition of parts (based on Trilobita)

Ecospace Comments

BODY SIZE: Based on component genera; 7 of 7 genera share 10 to < 100 length, 7 of 7 share 10 to < 100 width. LIFE HISTORY: Data from Meyer et al. (2009)
JA: carnivorous according to Fortey and Owens 1999

Taphonomy

low Mg calcite, chitin (based on Trilobita)

Primary Reference (PBDB)

W.T. Dean. 1983. Additional faunal data for the Bedinan Formation (Ordovician) of south-eastern Turkey. Special Papers in Palaeontology 30:89-105

Wikipedia

https://en.wikipedia.org/wiki/Calymenidae

Fossil Distribution

Subtaxa

Name	Status	Fossil Occurrences	Oldest	Youngest
Calymeninae ✝ subfamily	listed (PBDB)	557	485 Ma Cambrian	411 Ma Early/Lower Devonian
Flexicalymeninae ✝ subfamily	listed (PBDB)	805	470 Ma Middle Ordovician	419 Ma Early/Lower Devonian
Pharostomatinae ✝ subfamily	listed (PBDB)	108	485 Ma Cambrian	445 Ma Late/Upper Ordovician
Reedocalymeninae ✝ subfamily	listed (PBDB)	26	485 Ma Cambrian	453 Ma Late/Upper Ordovician
Alcymene ✝ genus	accepted (GBIF) listed (PBDB)
Apocalymene ✝ genus	doubtful (GBIF) listed (PBDB)	36	433 Ma Silurian	411 Ma Early/Lower Devonian
Arcticalymene ✝ genus	accepted (GBIF) listed (PBDB)	16	433 Ma Silurian	433 Ma Silurian
Calymene ✝ genus	accepted (GBIF) listed (PBDB)	472	485 Ma Cambrian	411 Ma Early/Lower Devonian
Calymenesun ✝ genus	accepted (GBIF) listed (PBDB)	18	479 Ma Early/Lower Ordovician	450 Ma Late/Upper Ordovician
Diacalymene ✝ genus	accepted (GBIF) listed (PBDB)	67	450 Ma Late/Upper Ordovician	426 Ma Silurian
Flexicalymene ✝ genus	accepted (GBIF) listed (PBDB)	789	470 Ma Middle Ordovician	433 Ma Silurian
Gravicalymene ✝ genus	accepted (GBIF) listed (PBDB)	103	466 Ma Middle Ordovician	408 Ma Early/Lower Devonian
Linguocalymene ✝ genus	doubtful (GBIF) listed (PBDB)	2	423 Ma Silurian	423 Ma Silurian
Liocalymene ✝ genus	doubtful (GBIF) listed (PBDB)	21	443 Ma Silurian	433 Ma Silurian
Neseuretus ✝ genus	accepted (GBIF) listed (PBDB)	133	485 Ma Cambrian	450 Ma Late/Upper Ordovician
Onnicalymene ✝ genus	accepted (GBIF) listed (PBDB)	7	456 Ma Late/Upper Ordovician	456 Ma Late/Upper Ordovician
Papillicalymene ✝ genus	accepted (GBIF) listed (PBDB)	9	450 Ma Late/Upper Ordovician	423 Ma Silurian
Pharostoma ✝ genus	accepted (GBIF) listed (PBDB)	39	467 Ma Middle Ordovician	445 Ma Late/Upper Ordovician
Platycalymene ✝ genus	accepted (GBIF) listed (PBDB)	25	466 Ma Middle Ordovician	458 Ma Late/Upper Ordovician
Pradoella ✝ genus	accepted (GBIF) listed (PBDB)	8	479 Ma Early/Lower Ordovician	466 Ma Middle Ordovician
Prionocheilus ✝ genus	accepted (GBIF) listed (PBDB)	66	485 Ma Cambrian	450 Ma Late/Upper Ordovician
Protocalymene ✝ genus	accepted (GBIF) listed (PBDB)	11	472 Ma Early/Lower Ordovician	466 Ma Middle Ordovician
Reacalymene ✝ genus	doubtful (GBIF) listed (PBDB)	1	461 Ma Middle Ordovician	461 Ma Middle Ordovician
Reedocalymene ✝ genus	accepted (GBIF) listed (PBDB)	6	479 Ma Early/Lower Ordovician	453 Ma Late/Upper Ordovician
Salterocoryphe ✝ genus	accepted (GBIF) listed (PBDB)	25	485 Ma Cambrian	461 Ma Middle Ordovician
Spathacalymene ✝ genus	doubtful (GBIF) listed (PBDB)
Sthenarocalymene ✝ genus	accepted (GBIF) listed (PBDB)	11	467 Ma Middle Ordovician	419 Ma Early/Lower Devonian
Synhomalonotus ✝ genus	accepted (GBIF) listed (PBDB)	11	479 Ma Early/Lower Ordovician	450 Ma Late/Upper Ordovician
Tapinocalymene ✝ genus	doubtful (GBIF) listed (PBDB)	1	433 Ma Silurian	433 Ma Silurian
Thelecalymene ✝ genus	accepted (GBIF) listed (PBDB)	2	450 Ma Late/Upper Ordovician	450 Ma Late/Upper Ordovician
Vietnamia ✝ genus	accepted (GBIF) listed (PBDB)	10	461 Ma Middle Ordovician	453 Ma Late/Upper Ordovician
Xuanenia ✝ genus	accepted (GBIF) listed (PBDB)	3	453 Ma Late/Upper Ordovician	453 Ma Late/Upper Ordovician

Synonymy List

Year	Name and Author
1840	Calymenidae Milne-Edwards
1957	Calymenidae Harrington and Leanza
1968	Calymenidae Robison and Pantoja-Alor p. 783
1983	Calymenidae Dean
1997	Calymenidae Whittington et al. p. 300
2011	Calymenidae Adrain p. 106

References

Dean W.T. (1983) Additional faunal data for the Bedinan Formation (Ordovician) of south-eastern Turkey, Special Papers in Palaeontology 30, 89-105

Harrington H. J., Leanza A. F. (1957) Ordovician trilobites of Argentina, Department of Geology, University of Kansas Special Publication. Lawrence: University of Kansas Press 1, 1-276

Robison R. A., Pantoja-Alor J. (1968) Tremadocian trilobites from the Nochixtlan Region, Oaxaca, Mexico, Journal of Paleontology 42 3, 767-800

Whittington H. B., Chatterton B. D. E., et al (1997) Trilobita. Introduction, Order Agnostina, Order Redlichiida. Volume 1, Treatise on Invertebrate Paleontology O, 1-530

Adrain J. M. (2011) Class Trilobita Walch, 1771, Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness 3148, 104-109

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

extinct group - via The Interim Register of Marine and Nonmarine Genera

International Commission on Zoological Nomenclature. Opinion 1433 Calymene Brongniart in Brongniart & Desmarest, 1822 (Trilobita): conserved.Bulletin of Zoological Nomenclature 44(2), 132-133 (1987) - via Official Lists and Indexes of Names in Zoology

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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