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Varanopidae ✝

Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)Not linked to a GBIF record.
PaleoBioDB ID (PBDB)91792https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=91792
Rankfamily (PBDB)
Taxonomy (PBDB)Life : Animalia : Chordata : Osteichthyes : Cotylosauria : Varanopidae
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-TriploblasticaLankester 1877
-NephrozoaJondelius et al. 2002
-DeuterostomiaGrobben 1908
phylumChordataHaeckel 1847
subphylumVertebrataLamarck 1801
superclassGnathostomataGegenbauer 1874
-Osteichthyes
-SarcopterygiiRomer 1955
subclassDipnotetrapodomorpha
subclassTetrapodomorpha
-Tetrapoda
-ReptiliomorphaSäve-Söderbergh 1934
-AnthracosauriaSäve-Söderbergh 1934
-Batrachosauria
-Cotylosauria
-AmniotaHaeckel 1866
-SynapsidaOsborn 1903
suborderEupelycosauriaKemp 1982
familyVaranopidaeRomer and Price 1940
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Varanopidaefamilybelongs to Sphenacodontiastated without evidenceRomer and Price, 1940
Varanopidaefamilybelongs to Sphenacodontiastated with evidenceKuhn, 1946
Varanopidaefamilybelongs to Sphenacodontiastated without evidenceHaughton and Brink, 1954
Varanopidaefamilybelongs to Sphenacodontiastated with evidenceKuhn, 1966
Varanopidaefamilybelongs to Sphenacodontiastated without evidenceLangston and Reisz, 1981
Varanopidaefamilybelongs to Eupelycosauriastated without evidenceKemp, 1982
Varanopidaefamilybelongs to Pelycosauriastated with evidenceEberth and Brinkman, 1983
Varanopidaefamilybelongs to Eupelycosauriastated without evidenceReisz, 1986
Varanopidaefamilybelongs to PelycosauriaimpliedCarroll, 1988
Varanopidaefamilybelongs to Eupelycosauriastated without evidenceDilkes and Reisz, 1996
Varanopidaefamilybelongs to Eupelycosauriastated with evidenceReisz and Dilkes, 2003
Varanopidaefamilybelongs to Eupelycosauriastated without evidenceAnderson and Reisz, 2004
Varanopidaefamilybelongs to EupelycosauriaimpliedMartens et al., 2005
Varanopidaefamilybelongs to Eupelycosauriastated with evidenceMaddin et al., 2006
Varanopidaefamilybelongs to SynapsidaimpliedReisz and Modesto, 2007
Varanopidaefamilybelongs to Varanopoideastated without evidenceIvakhnenko, 2008
Varanopidaefamilybelongs to Eupelycosauriastated with evidenceEvans et al., 2009
Status (PBDB)extinct
Taxon Size (PBDB)39
First Recorded Appearance303 - 299 Ma
Carboniferous
Last Recorded Appearance265 - 260 Ma
Permian
Environmentterrestrial (based on Amniota)
Motilityactively mobile (based on Osteichthyes)
Taphonomyphosphatic (based on Vertebrata)
Primary Reference (PBDB)A. S. Romer and L. I. Price. 1940. Review of the Pelycosauria. Geological Society of America Special Paper 28:1-538

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Mycterosaurinae
subfamily
listed (PBDB)1290 Ma
Permian
290 Ma
Permian
Apsisaurus
genus
accepted (GBIF)
listed (PBDB)
1296 Ma
Permian
296 Ma
Permian
Archaeovenator
genus
accepted (GBIF)
listed (PBDB)
1303 Ma
Carboniferous
303 Ma
Carboniferous
Ascendonanus
genus
listed (PBDB)1296 Ma
Permian
296 Ma
Permian
Pyozia
genus
accepted (GBIF)
listed (PBDB)
1273 Ma
Permian
273 Ma
Permian
Neovaranopsia
unranked clade
listed (PBDB)31303 Ma
Carboniferous
265 Ma
Permian

Synonymy List

YearName and Author
1937Elliotsmithidae Broom
1940Varanopsidae Romer and Price pp. 18, 268
1946Varanopsidae Kuhn p. 56
1954Varanopsidae Haughton and Brink p. 5
1962Thrausmosaurus Fox p. 302 figs. 4-6
1962Thrausmosaurus serratidens Fox p. 302 figs. 4-6
1966Varanopsidae Kuhn p. 120
1981Varanopseidae Langston and Reisz p. 74
1982Varanopidae Kemp p. 348
1983Varanopseidae Eberth and Brinkman p. 2
1986Varanopseidae Reisz p. 63
1986Thrausmosaurus Reisz p. 85
1986Thrausmosaurus serratidens Reisz p. 85
YearName and Author
1988Thrausmosaurus Carroll
1988Varanopseidae Carroll
1996Varanopseidae Dilkes and Reisz p. 1167
2003Varanopidae Reisz and Dilkes p. 669
2004Varanopidae Anderson and Reisz p. 173
2004Varanopidae Reisz and Laurin p. 378
2005Varanopseidae Martens et al.
2006Varanopidae Maddin et al. p. 957
2007Varanopidae Reisz and Modesto p. 734
2008Varanopidae Ivakhnenko
2009Varanopidae Evans et al. p. 220
2019Varanopidae Maho et al.

References

Romer A. S., Price L. I. (1940) Review of the Pelycosauria, Geological Society of America Special Paper 28, 1-538
Kuhn O. (1946) Das System der fossilen und rezenten Amphibien und Reptilien [The system of fossil and recent amphibians and reptiles], Bericht der Naturforschenden Gesellschaft in Bamberg 29, 49-67
Haughton S. H., Brink A. S. (1954) A bibliographical list of Reptilia from the Karroo Beds of South Africa, Palaeontologia Africana 2, 1-187
Fox R. C. (1962) Two New Pelycosaurs from the Lower Permian of Oklahoma, University of Kansas Publications, Museum of Natural History 12 6, 297-307
Kuhn O. W. M. (1966) , Die Reptilien. System und Stammesgeschichte [The Reptiles. Systematics and Phylogeny.], 1-154
Langston W., Reisz R. R. (1981) Aerosaurus wellesi, new species, a varanopseid mammal-like reptile (Synapsida: Pelycosauria) from the Lower Permian of New Mexico, Journal of Vertebrate Paleontology 1 1, 73-96
Kemp T. S. (1982) , Mammal-like Reptiles and the Origin of Mammals, 1-363
Eberth D. A., Brinkman D. (1983) Ruthiromia elcobriensis, a New Pelycosaur from El Cobre Canyon, New Mexico, Breviora 474, 1-26
Reisz R. R. (1986) Pelycosauria, Handbuch der Paläoherpetologie / Encyclopedia of Paleoherpetology 17A, 1-102
Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698
Dilkes D. W., Reisz R. R. (1996) First record of a basal synapsid (‘mammal-like reptile’) in Gondwana, Proceedings of the Royal Society of London, Series B 263, 1165-1170
Reisz R. R., Dilkes D. W. (2003) Archaeovenator hamiltonensis, a new varanopid (Synapsida: Eupelycosauria) from the Upper Carboniferous of Kansas, Canadian Journal of Earth Sciences 40 4, 667-678
Anderson J. S., Reisz R. R. (2004) Pyozia mesenensis, a New, Small Vapanopid (Synapsida: Eupelycosauria) from Russia: ‘‘Pelycosaur’’ Diversity in the Middle Permian, Journal of Vertebrate Paleontology 24 1, 173-179
Reisz R. R., Laurin M. (2004) A reevaluation of the enigmatic Permian synapsid Watongia and of its stratigraphic significance, Canadian Journal of Earth Sciences 41 4, 377-386
Martens T., Berman D. S., et al (2005) The Bromacker Quarry - the Most Important Locality of Lower Permian Terrestrial Vertebrate Fossils Outside of North America, The Nonmarine Permian. New Mexico Museum of Natural History and Science Bulletin 30, 214-215
Maddin H. C., Evans D. C., et al (2006) An Early Permian varanodontine varanopid (Synapsida: Eupelycosauria) from the Richards Spur locality, Oklahoma, Journal of Vertebrate Paleontology 26 4, 957-966 doi:10.1671/0272-4634(2006)26[957:aepvvs]2.0.co;2
Reisz R. R., Modesto S. P. (2007) Heleosaurus scholtzi from the Permian of South Africa: a Varanopid Synapsid, not a Diapsid Reptile, Journal of Vertebrate Paleontology 27 3, 734-739
Ivakhnenko M. F. (2008) Podklass Ophiacomorpha, Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast' 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], GEOS, Moscow, 95-100
Evans D. C., Maddin H. C., et al (2009) A Re-Evaluation of Sphenacodontid Synapsid Material from the Lower Permian Fissure Fills near Richards Spur, Oklahoma, Palaeontology 52 1, 219-227 doi:10.1111/j.1475-4983.2008.00837.x
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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