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Athyrididae ✝

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Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Composita subtilita

Kohl’s Ranch fossil locality, Gila County, Arizona, USA

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3252335	https://www.gbif.org/species/3252335
PaleoBioDB ID (PBDB)	81197	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=81197

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Brachiopoda : Rhynchonellata : Athyridida : Athyrididae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Spiralia
superphylum	Lophotrochozoa
-	Lophophorata
phylum	Brachiopoda	Cuvier 1805
subphylum	Rhynchonelliformea	Williams et al. 1996
class	Rhynchonellata	Williams et al. 1996
order	Athyridida	Boucot et al. 1964
suborder	Athyrididina	Boucot et al. 1964
superfamily	Athyridoidea	McCoy 1844
family	Athyrididae	McCoy 1841

Scientific Name

Athyrididae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Athyrididae	family	belongs to Rostrospiracea	stated without evidence	Hoare, 1961
Athyrididae	family	belongs to Athyridacea	stated with evidence	Rubel and Modzalevskaya, 1967
Athyrididae	family	belongs to Athyridacea	stated without evidence	Bassett and Cocks, 1974
Athyrididae	family	belongs to Athyridoidea	stated without evidence	Savage and Baxter, 1995
Athyrididae	family	belongs to Athyroidea	stated with evidence	Jin and Chatterton, 1997
Athyrididae	family	belongs to Athyridoidea	stated with evidence	Williams et al., 2002
Athyrididae	family	belongs to Athyridoidea	stated with evidence	Musteikis and Modzalevskaya, 2002
Athyrididae	family	misspelling of Athyrididae	stated with evidence	Williams et al., 2002
Athyrididae	family	belongs to Athyridioidea	stated without evidence	Shen et al., 2003

Status (PBDB)

extinct

Taxon Size (PBDB)

367

First Recorded Appearance

443 - 439 Ma
Silurian

Last Recorded Appearance

183 - 182 Ma
Early Jurassic

Motility

stationary (based on Athyrididina)

Vision

blind (based on Rhynchonellata)

Diet

suspension feeder (based on Athyrididina)

Taphonomy

low Mg calcite (based on Brachiopoda)

Primary Reference (PBDB)

J. Phillips. 1841. Figures and descriptions of the Palaeozoic fossils of Cornwall, Devon, and West Somerset; observed in the course of the Ordinance Geological Survey of that district. Geological Society of London

Fossil Distribution

Subtaxa

Name	Status	Fossil Occurrences	Oldest	Youngest
Athyridinae ✝ subfamily	listed (PBDB)	1,148	443 Ma Silurian	228 Ma Late/Upper Triassic
Athyrisininae ✝ subfamily	listed (PBDB)	51	427 Ma Silurian	345 Ma Carboniferous
Cleiothyrididinae ✝ subfamily	listed (PBDB)	1,841	388 Ma Middle Devonian	254 Ma Permian
Comelicaniinae ✝ subfamily	listed (PBDB)	33	254 Ma Permian	254 Ma Permian
Didymothyridinae ✝ subfamily	listed (PBDB)	178	441 Ma Silurian	388 Ma Middle Devonian
Helenathyridinae ✝ subfamily	listed (PBDB)	1	383 Ma Late/Upper Devonian	383 Ma Late/Upper Devonian
Janicepsinae ✝ subfamily	listed (PBDB)	45	260 Ma Permian	254 Ma Permian
Lochengiinae ✝ subfamily	listed (PBDB)	47	372 Ma Late/Upper Devonian	265 Ma Permian
Plicathyridinae ✝ subfamily	listed (PBDB)	151	412 Ma Early/Lower Devonian	380 Ma Late/Upper Devonian
Pradoiinae ✝ subfamily	listed (PBDB)	2	419 Ma Early/Lower Devonian	403 Ma Early/Lower Devonian
Protathyridinae ✝ subfamily	listed (PBDB)
Spirigerellinae ✝ subfamily	listed (PBDB)	3,398	411 Ma Early/Lower Devonian	222 Ma Late/Upper Triassic
Transcaucasathyriinae ✝ subfamily	listed (PBDB)	323	290 Ma Permian	252 Ma Permian
Xenosariinae ✝ subfamily	listed (PBDB)	5	269 Ma Permian	260 Ma Permian
Actinoconchus ✝ genus	accepted (GBIF) listed (PBDB)	39	376 Ma Late/Upper Devonian	260 Ma Permian
Anathyris ✝ genus	accepted (GBIF) listed (PBDB)	52	412 Ma Early/Lower Devonian	380 Ma Late/Upper Devonian
Araxathyris ✝ genus	accepted (GBIF) listed (PBDB)	355	279 Ma Permian	222 Ma Late/Upper Triassic
Athyris ✝ genus	accepted (GBIF) listed (PBDB)	940	443 Ma Silurian	228 Ma Late/Upper Triassic
Bajtugania ✝ genus	accepted (GBIF) listed (PBDB)	52	272 Ma Permian	260 Ma Permian
Baliqliqia ✝ genus	accepted (GBIF) listed (PBDB)	9	290 Ma Permian	279 Ma Permian
Buchanathyris ✝ genus	accepted (GBIF) listed (PBDB)	15	419 Ma Early/Lower Devonian	408 Ma Early/Lower Devonian
Cleiothyridina ✝ genus	accepted (GBIF) listed (PBDB)	1,701	388 Ma Middle Devonian	254 Ma Permian
Collarothyris ✝ genus	accepted (GBIF) listed (PBDB)	21	426 Ma Silurian	423 Ma Silurian
Comelicania ✝ genus	accepted (GBIF) listed (PBDB)	27	254 Ma Permian	254 Ma Permian
Comelicothyris ✝ genus	accepted (GBIF) listed (PBDB)	13	254 Ma Permian	254 Ma Permian
Composita ✝ genus	accepted (GBIF) listed (PBDB)	2,002	380 Ma Late/Upper Devonian	254 Ma Permian
Crinisarina ✝ genus	accepted (GBIF) listed (PBDB)	23	383 Ma Late/Upper Devonian	359 Ma Late/Upper Devonian
Cryptospirifer ✝ genus	accepted (GBIF) listed (PBDB)	32	372 Ma Late/Upper Devonian	265 Ma Permian
Didymothyris ✝ genus	accepted (GBIF) listed (PBDB)	47	441 Ma Silurian	423 Ma Silurian
Glassina ✝ genus	accepted (GBIF) listed (PBDB)	9	433 Ma Silurian	423 Ma Silurian
Gruntallina ✝ genus	accepted (GBIF) listed (PBDB)	6	254 Ma Permian	254 Ma Permian
Gruntea ✝ genus	accepted (GBIF) listed (PBDB)	10	260 Ma Permian	254 Ma Permian
Himathyris ✝ genus	accepted (GBIF) listed (PBDB)	11	290 Ma Permian	260 Ma Permian
Janiceps ✝ genus	accepted (GBIF) listed (PBDB)	32	260 Ma Permian	254 Ma Permian
Johnsonathyris ✝ genus	accepted (GBIF) listed (PBDB)	3	372 Ma Late/Upper Devonian	372 Ma Late/Upper Devonian
Juxathyris ✝ genus	accepted (GBIF) listed (PBDB)	152	290 Ma Permian	252 Ma Permian
Kintathyris ✝ genus	accepted (GBIF) listed (PBDB)	1	296 Ma Permian	296 Ma Permian
Leiothycridina ✝ genus	accepted (GBIF) listed (PBDB)	6	359 Ma Late/Upper Devonian	269 Ma Permian
Leptathyris ✝ genus	accepted (GBIF) listed (PBDB)	38	409 Ma Early/Lower Devonian	388 Ma Middle Devonian
Mayangella ✝ genus	accepted (GBIF) listed (PBDB)	3	260 Ma Permian	260 Ma Permian
Nordathyris ✝ genus	accepted (GBIF) listed (PBDB)
Pachyplax ✝ genus	accepted (GBIF) listed (PBDB)	28	408 Ma Early/Lower Devonian	403 Ma Early/Lower Devonian
Pachyplaxoides ✝ genus	doubtful (GBIF) listed (PBDB)	1	380 Ma Late/Upper Devonian	380 Ma Late/Upper Devonian
Pinegathyris ✝ genus	accepted (GBIF) listed (PBDB)	44	299 Ma Carboniferous	260 Ma Permian
Planalvus ✝ genus	accepted (GBIF) listed (PBDB)	8	411 Ma Early/Lower Devonian	345 Ma Carboniferous
Posicomta ✝ genus	accepted (GBIF) listed (PBDB)	16	279 Ma Permian	272 Ma Permian
Pradoia ✝ genus	accepted (GBIF) listed (PBDB)	1	403 Ma Early/Lower Devonian	403 Ma Early/Lower Devonian
Protathyris ✝ genus	accepted (GBIF) listed (PBDB)	105	443 Ma Silurian	409 Ma Early/Lower Devonian
Pseudoprotathyris ✝ genus	accepted (GBIF) listed (PBDB)	37	441 Ma Silurian	419 Ma Early/Lower Devonian
Rectambitus ✝ genus	accepted (GBIF) listed (PBDB)	7	260 Ma Permian	254 Ma Permian
Septospirigerella ✝ genus	accepted (GBIF) listed (PBDB)	74	269 Ma Permian	222 Ma Late/Upper Triassic
Spirigera ✝ genus	accepted (GBIF) listed (PBDB)	50	272 Ma Permian	209 Ma Late/Upper Triassic
Spirigerella ✝ genus	accepted (GBIF) listed (PBDB)	912	411 Ma Early/Lower Devonian	254 Ma Permian
Tarimathyris ✝ genus	accepted (GBIF) listed (PBDB)	30	290 Ma Permian	254 Ma Permian
Titanothyris ✝ genus	accepted (GBIF) listed (PBDB)	14	279 Ma Permian	269 Ma Permian
Tongzithyris ✝ genus	accepted (GBIF) listed (PBDB)	10	265 Ma Permian	254 Ma Permian
Transcaucasathyris ✝ genus	accepted (GBIF) listed (PBDB)	171	265 Ma Permian	254 Ma Permian
Xenosaria ✝ genus	accepted (GBIF) listed (PBDB)	5	269 Ma Permian	260 Ma Permian

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Athyris naliensis Reed, 1944	GBIF	species	Animalia : Brachiopoda : Rhynchonellata : Athyridida : Athyrididae : *Athyris naliensis*
Athyridae	GBIF	family	Animalia : Brachiopoda : Rhynchonellata : Athyridida : Athyridae

Synonymy List

Year	Name and Author
1841	Athyrididae McCoy p. 94
1944	Athyris naliensis Reed p. 532
1961	Athyridae Hoare p. 89
1967	Athyrididae Rubel and Modzalevskaya
1974	Athyrididae Bassett and Cocks
1974	Athyrididae Brunton and Champion p. 828

Year	Name and Author
1978	Athyrididae Savage et al. p. 381
1995	Athyrididae Savage and Baxter p. 1039
1997	Athyrididae Jin and Chatterton
2002	Athyrididae Musteikis and Modzalevskaya
2002	Athyrididae Williams et al.
2003	Athyrididae Shen et al. p. 248

References

Phillips J. Figures and descriptions of the Palaeozoic fossils of Cornwall, Devon, and West Somerset; observed in the course of the Ordinance Geological Survey of that district, Geological Society of London, 1-231

Reed F. R. C. (1944) Brachiopoda and Mollusca from the Productus limestones of the Salt Range, Palaeontogica Indica, Memoirs of the Geological Survey of India, New Series 23 2, 1-596

Hoare R. D. (1961) Desmoinesian Brachiopoda and Mollusca from southwest Missouri, Missouri University Studies 36, 1-262

Rubel M., Modzalevskaya T. (1967) New Siluria Brachiopods of the family Athyrididae, Proceedings of the Estonian Academy of Sciences 3

Bassett M. G., Cocks L. R. M. (1974) A review of Silurian brachiopods from Gotland, Fossils and Strata 3, 1-56

Brunton C. H. C., Champion C. (1974) A Lower Carboniferous brachiopod fauna from the Manifold Valley, Staffordshire, Palaeontology 17 4, 811-840

Savage N. M., Eberlein G. D., et al (1978) Upper Devonian brachiopods from the Port Refugio Formation, Suemez Island, southeastern Alaska, Journal of Paleontology 52 2, 370-393

Savage N. M., Baxter M. (1995) Late Devonian (Frasnian) brachiopods from the Wadleigh Limestone, Southeastern Alaska, Journal of Paleontology 69 6, 1029-1046

Jin J., Chatterton B. D. E. (1997) Latest Ordovician-Silurian articulate brachiopods and biostratigraphy of the Avalanche Lake area, southwestern District of Mackenzie, Canada, Palaeontographica Canadiana 13, 1-62

Musteikis P., Modzalevskaya T. L. (2002) Some Silurian Brachiopods from Lithuania and their Palaeobiogeographical significance, Palaeontology 45 3, 595-626

Williams A., Brunton C. H. C., et al (2002) Part H Brachiopoda (revised): Volume 4, Rhynchonelliformea (part), Treatise on Invertebrate Paleontology, 921-1688

Shen S. Z., Shi G. R., et al (2003) A Wuchiapingian (Late Permian) brachiopod fauna from an exotic block in the Indus-Tsangpo Suture Zone, southern Tibet, and its palaeobiogeographical and tectonic implications, Palaeontology 46 2, 225-256

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Museum Victoria KEmu database (Oct 2006) - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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