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Coptoclavidae ✝

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Description

Coptoclavidae is an extinct family of beetles in the suborder Adephaga. The Coptoclavidae lived from the Jurassicto the Early Cretaceous. Coptoclavidae is a clade of dytiscoidea. The increase in the teleost fish population and the competition of gyrinidae and dytiscidae is believed to have caused their extinction.

From Wikipedia article at https://en.wikipedia.org/wiki/Coptoclavidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3263914	https://www.gbif.org/species/3263914
PaleoBioDB ID (PBDB)	153821	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=153821

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Coptoclavidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Coptoclavidae	Ponomarenko 1961

Scientific Name

Coptoclavidae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Coptoclavidae	family	belongs to Adephaga	stated without evidence	Carpenter, 1992
Coptoclavidae	family	belongs to Adephaga	stated with evidence	Soriano et al., 2007

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

252 - 247 Ma
Phanerozoic

Last Recorded Appearance

113 - 101 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

F. M. Carpenter. 1992. Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4:1-655

Wikipedia

https://en.wikipedia.org/wiki/Coptoclavidae

Fossil Distribution

Subtaxa

Name	Status	Fossil Occurrences	Oldest	Youngest
Charonoscaphinae ✝ subfamily	listed (PBDB)	9	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Coptoclavinae ✝ subfamily	listed (PBDB)	37	164 Ma Middle Jurassic	113 Ma Early/Lower Cretaceous
Coptoclaviscinae ✝ subfamily	listed (PBDB)	21	247 Ma Early/Lower Triassic	125 Ma Early/Lower Cretaceous
Hispanoclavinae ✝ subfamily	listed (PBDB)	2	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Timarchopsinae ✝ subfamily	listed (PBDB)	64	232 Ma Late/Upper Triassic	125 Ma Early/Lower Cretaceous
Agrascapha ✝ genus	accepted (GBIF) listed (PBDB)	1	237 Ma Middle Triassic	237 Ma Middle Triassic
Amblycephalonius ✝ genus	accepted (GBIF) listed (PBDB)	3	183 Ma Early Jurassic	183 Ma Early Jurassic
Bolbonectes ✝ genus	accepted (GBIF) listed (PBDB)	5	164 Ma Middle Jurassic	129 Ma Early/Lower Cretaceous
Charonoscapha ✝ genus	accepted (GBIF) listed (PBDB)	6	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Charonoscaphidia ✝ genus	accepted (GBIF) listed (PBDB)	3	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Coptoclavella ✝ genus	accepted (GBIF) listed (PBDB)	15	166 Ma Middle Jurassic	125 Ma Early/Lower Cretaceous
Coptoclavia ✝ genus	accepted (GBIF) listed (PBDB)	1	252 Ma Permian	252 Ma Permian
Coptoclavisca ✝ genus	accepted (GBIF) listed (PBDB)	2	140 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Daohugounectes ✝ genus	accepted (GBIF) listed (PBDB)	4	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Ditomoptera ✝ genus	accepted (GBIF) listed (PBDB)	11	151 Ma Late Jurassic	151 Ma Late Jurassic
Euroscapha ✝ genus	accepted (GBIF) listed (PBDB)	2	237 Ma Middle Triassic	237 Ma Middle Triassic
Exedia ✝ genus	accepted (GBIF) listed (PBDB)	2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Hispanoclavina ✝ genus	accepted (GBIF) listed (PBDB)	2	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Holcoptera ✝ genus	accepted (GBIF) listed (PBDB)	37	228 Ma Late/Upper Triassic	197 Ma Early Jurassic
Hoyaclava ✝ genus	accepted (GBIF) listed (PBDB)	1	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Megacoptoclava ✝ genus	accepted (GBIF) listed (PBDB)	1	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Ovonectes ✝ genus	accepted (GBIF) listed (PBDB)	1	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Protonectes ✝ genus	accepted (GBIF) listed (PBDB)	1	232 Ma Late/Upper Triassic	232 Ma Late/Upper Triassic
Pseudohydrophilus ✝ genus	accepted (GBIF) listed (PBDB)	8	151 Ma Late Jurassic	151 Ma Late Jurassic
Stargelytron ✝ genus	accepted (GBIF) listed (PBDB)	4	247 Ma Early/Lower Triassic	232 Ma Late/Upper Triassic
Stygeonectes ✝ genus	accepted (GBIF) listed (PBDB)	17	183 Ma Early Jurassic	125 Ma Early/Lower Cretaceous
Timarchopsis ✝ genus	accepted (GBIF) listed (PBDB)	12	191 Ma Early Jurassic	133 Ma Early/Lower Cretaceous
Tuhanectes ✝ genus	accepted (GBIF) listed (PBDB)	1	168 Ma Middle Jurassic	168 Ma Middle Jurassic

Synonymy List

Year	Name and Author
1961	Coptoclavidae Ponomarenko p. 68
1992	Coptoclavidae Carpenter
2007	Coptoclavidae Soriano et al. p. 527
2011	Coptoclavidae Bouchard et al.

References

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655

Soriano C., Ponomarenko A. G., et al (2007) Coptoclavid beetles (Coleoptera: Adephaga) from the Lower Cretaceous of Spain: A new feeding strategy in beetles, Palaeontology 50 2, 525-536

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Slipinski, A.; Smith, A. (2011). Family-group names in Coleoptera (Insecta). ZooKeys. 88: 1-972. - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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