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Acroceridae

Description

The Acroceridae are a small family of odd-looking flies. They have a hump-backed appearance with a strikingly small head, generally with a long proboscis for accessing nectar. They are rare and not widely known. The most frequently applied common names are small-headed flies or hunch-back flies. Many are bee or wasp mimics. Because they are parasitoids of spiders, they also are sometimes known as spider flies.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)3328https://www.gbif.org/species/3328
PaleoBioDB ID (PBDB)138867https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=138867
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Diptera : Acroceridae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-Holometabola
-Aparaglossata
-Mecopterida
superorderAntliophora
orderDipteraLinnaeus 1758
-NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
-Homeodactyla
superfamilyNemestrinoideaMacquart 1834
familyAcroceridaeLeach 1815
Common Namespider fly
Scientific NameAcroceridae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Acroceridaefamilybelongs to Brachycerastated without evidenceCarpenter, 1992
Acroceridaefamilybelongs to Nemestrinoideastated with evidenceGrimaldi et al., 2011
Status (PBDB)extant
Taxon Size (PBDB)31
Extant Size (PBDB)7 (23%)
First Recorded Appearance166 - 157 Ma
Jurassic
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)F. M. Carpenter. 1992. Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4:1-655
Common Name(s) Spinvliegen, Small-headed Flies, Kulflugor, Spider Fly
Wikipediahttps://en.wikipedia.org/wiki/Acroceridae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Acrocerinae
subfamily
listed (PBDB)299.6 Ma
Late/Upper Cretaceous
0 Ma
Extant
Archocyrtinae
subfamily
listed (PBDB)2166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Cyrtinae
subfamily
listed (PBDB)338.0 Ma
Eocene
0 Ma
Extant
Ogcodinae
subfamily
listed (PBDB)438.0 Ma
Eocene
0 Ma
Extant
Philopotinae
subfamily
listed (PBDB)938.0 Ma
Eocene
0 Ma
Extant
Acrocera
genus
accepted (GBIF)
listed (PBDB)
0 Ma
Extant
Africaterphis
genus
accepted (GBIF)No associated record in PBDB
Apelleia
genus
accepted (GBIF)No associated record in PBDB
Apsona
genus
accepted (GBIF)No associated record in PBDB
Archaeterphis
genus
accepted (GBIF)
listed (PBDB)
338.0 Ma
Eocene
38.0 Ma
Eocene
Archipialea
genus
accepted (GBIF)No associated record in PBDB
Archocyrtus
genus
accepted (GBIF)
listed (PBDB)
2166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Asopsebius
genus
accepted (GBIF)No associated record in PBDB
Astomella
genus
accepted (GBIF)No associated record in PBDB
Astomelloides
genus
accepted (GBIF)No associated record in PBDB
Burmacyrtus
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Camposella
genus
accepted (GBIF)No associated record in PBDB
Coquena
genus
accepted (GBIF)No associated record in PBDB
Corononcodes
genus
accepted (GBIF)No associated record in PBDB
Cyrtus
genus
accepted (GBIF)No associated record in PBDB
Dimacrocolus
genus
accepted (GBIF)No associated record in PBDB
Eulonchiella
genus
accepted (GBIF)
listed (PBDB)
338.0 Ma
Eocene
38.0 Ma
Eocene
Eulonchus
genus
accepted (GBIF)No associated record in PBDB
Exetasis
genus
accepted (GBIF)No associated record in PBDB
Glaesoncodes
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Hadrogaster
genus
accepted (GBIF)No associated record in PBDB
Helle
genus
accepted (GBIF)No associated record in PBDB
Holops
genus
accepted (GBIF)No associated record in PBDB
Lasia
genus
accepted (GBIF)No associated record in PBDB
Lasioides
genus
accepted (GBIF)No associated record in PBDB
Leucopsina
genus
accepted (GBIF)No associated record in PBDB
Megalybus
genus
accepted (GBIF)No associated record in PBDB
Meruia
genus
accepted (GBIF)No associated record in PBDB
Mesophysa
genus
accepted (GBIF)No associated record in PBDB
Neolasia
genus
accepted (GBIF)No associated record in PBDB
Neopanops
genus
accepted (GBIF)No associated record in PBDB
Neophilopota
genus
accepted (GBIF)No associated record in PBDB
Nipponcyrtus
genus
accepted (GBIF)No associated record in PBDB
Ocnaea
genus
accepted (GBIF)No associated record in PBDB
Ogcodes
genus
accepted (GBIF)
listed (PBDB)
320.4 Ma
Miocene
0 Ma
Extant
Oligoneura
genus
accepted (GBIF)
listed (PBDB)
0 Ma
Extant
Oncodes
genus
accepted (GBIF)No associated record in PBDB
Opsebius
genus
accepted (GBIF)No associated record in PBDB
Panocalda
genus
accepted (GBIF)No associated record in PBDB
Panops
genus
accepted (GBIF)No associated record in PBDB
Paracyrtus
genus
accepted (GBIF)No associated record in PBDB
Parahelle
genus
accepted (GBIF)No associated record in PBDB
Philopota
genus
accepted (GBIF)No associated record in PBDB
Physegastrella
genus
accepted (GBIF)No associated record in PBDB
Pialea
genus
accepted (GBIF)No associated record in PBDB
Prophilopota
genus
accepted (GBIF)
listed (PBDB)
238.0 Ma
Eocene
38.0 Ma
Eocene
Psilodera
genus
accepted (GBIF)No associated record in PBDB
Pterodontia
genus
accepted (GBIF)No associated record in PBDB
Pteropexus
genus
accepted (GBIF)No associated record in PBDB
Quasi
genus
accepted (GBIF)No associated record in PBDB
Rhysogaster
genus
accepted (GBIF)No associated record in PBDB
Sabroskya
genus
accepted (GBIF)No associated record in PBDB
Schlingeriella
genus
accepted (GBIF)No associated record in PBDB
Schlingeromyia
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Sphaerops
genus
accepted (GBIF)No associated record in PBDB
Stenopialea
genus
accepted (GBIF)No associated record in PBDB
Subcyrtus
genus
accepted (GBIF)No associated record in PBDB
Terphis
genus
accepted (GBIF)No associated record in PBDB
Thyllis
genus
accepted (GBIF)No associated record in PBDB
Turbopsebius
genus
accepted (GBIF)No associated record in PBDB
Villalites
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Villalus
genus
accepted (GBIF)No associated record in PBDB
Pterondontia misella
species
accepted (GBIF)No associated record in PBDB

Synonymy List

YearName and Author
1815Acroceridae Leach p. 162
1992Acroceridae Carpenter
2011Acroceridae Grimaldi et al. p. 306
2018Acroceridae Shin et al.

References

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655
Grimaldi D. A., Arillo A., et al (2011) Brachyceran Diptera (Insecta) in Cretaceous ambers, Part IV, significant new Orthorrhaphous Taxa, ZooKeys 148, 293-332
Shin S., Bayless K. M., et al (2018) Taxon sampling to address an ancient rapid radiation: a supermatrix phylogeny of early brachyceran flies (Diptera), Systematic Entomology doi:10.1111/syen.12275
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
Pape, T.; Blagoderov, B.; Mostovski, M. B. (2011). Order Diptera Linnaeus, 1758. In Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa. 3148: 222-229. - via The Interim Register of Marine and Nonmarine Genera
Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas
Systema Dipterorum (2013) http://www.diptera.org/About.php - via Dyntaxa. Svensk taxonomisk databas
Shewell, G. E.,Wood, D. M.,Vockeroth, J. R.,Peterson, B. V.,Schlinger, E. I.,Teskey, H. J.,McAlpine, J. F. (1981) Chapter 43. Acroceridae - via The Diptera Site
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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