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Apsilocephalidae

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Description

The Apsilocephalidae comprise a family of flies in the superfamily Asiloidea. The family was proposed in 1991.

From Wikipedia article at https://en.wikipedia.org/wiki/Apsilocephalidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3331	https://www.gbif.org/species/3331
PaleoBioDB ID (PBDB)	193278	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=193278

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Diptera : Apsilocephalidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Mecopterida
superorder	Antliophora
order	Diptera	Linnaeus 1758
-	Neodiptera	Michelsen 1994
suborder	Brachycera	Zetterstedt 1842
-	Heterodactyla
superfamily	Asiloidea	Latreille 1802
family	Apsilocephalidae	Nagatomi et al. 1991

Scientific Name

Apsilocephalidae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Apsilocephalidae	family	belongs to Brachycera	stated with evidence	Nagatomi et al., 1991
Apsilocephalidae	family	subjective synonym of Rhagionempididae	stated with evidence	Nagatomi and Yang, 1998
Apsilocephalidae	family	belongs to Brachycera	stated with evidence	Hauser and Irwin, 2005
Apsilocephalidae	family	belongs to Asiloidea	stated with evidence	Grimaldi et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

2 (18%)

First Recorded Appearance

99.6 - 93.5 Ma
Late/Upper Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

A. Nagatomi, T. Saigusa, and H. Nagatomi, L. Lyneborg. 1991. Apsilocephalidae, a new family of the orthorrhaphous Brachycera (Insecta, Diptera). Zoological Science 8:579-591

Wikipedia

https://en.wikipedia.org/wiki/Apsilocephalidae

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Apsilocephala genus	accepted (GBIF) listed (PBDB)		3	38.0 Ma Eocene	0 Ma Extant
Burmapsilocephala ✝ genus	accepted (GBIF) listed (PBDB)		2	99.6 Ma Late/Upper Cretaceous	99.6 Ma Late/Upper Cretaceous
Clesthentia genus	accepted (GBIF)	No associated record in PBDB
Clesthentiella genus	accepted (GBIF)	No associated record in PBDB
Irwinimyia ✝ genus	accepted (GBIF) listed (PBDB)		1	99.6 Ma Late/Upper Cretaceous	99.6 Ma Late/Upper Cretaceous
Kaurimyia genus	accepted (GBIF)	No associated record in PBDB
Kumaromyia ✝ genus	accepted (GBIF) listed (PBDB)		1	99.6 Ma Late/Upper Cretaceous	99.6 Ma Late/Upper Cretaceous
Myanmarpsilocephala ✝ genus	accepted (GBIF) listed (PBDB)		1	99.6 Ma Late/Upper Cretaceous	99.6 Ma Late/Upper Cretaceous

Synonymy List

Year	Name and Author
1991	Apsilocephalidae Nagatomi et al. p. 579
2005	Apsilocephalidae Hauser and Irwin p. 397
2011	Apsilocephalidae Grimaldi et al. p. 317
2011	Apsilocephalidae Wiegmann et al.
2015	Apsilocephalidae Winterton and Ware
2016	Apsilocephalidae Grimaldi p. 76
2016	Apsilocephalidae Winterton et al.
2018	Apsilocephalidae Shin et al.

References

Hauser M., Irwin M. E. (2005) Fossil Therevidae (Insecta: Diptera) from Florissant, Colorado (upper Eocene), Journal of Systematic Palaeontology 3 4, 393-401

Grimaldi D. A., Arillo A., et al (2011) Brachyceran Diptera (Insecta) in Cretaceous ambers, Part IV, significant new Orthorrhaphous Taxa, ZooKeys 148, 293-332

Wiegmann B. M., Trautwein M. D., et al (2011) Episodic radiations in the fly tree of life, Proceedings of the National Academy of Sciences 108, 5690-5695

Winterton S. L., Ware J. L. (2015) Phylogeny, divergence times and biogeography of window flies (Scenopinidae) and the therevoid clade (Diptera: Asiloidea), Systematic Entomology 40, 491-519 doi:10.1111/syen.12117

Grimaldi D. A. (2016) Diverse orthorrhaphan flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII, Bulletin of the American Museum of Natural History 408, 1-131

Winterton S. L., Hardy N. B., et al (2016) The phylogeny of stiletto flies (Diptera: Therevidae), Systematic Entomology 41, 144-161

Shin S., Bayless K. M., et al (2018) Taxon sampling to address an ancient rapid radiation: a supermatrix phylogeny of early brachyceran flies (Diptera), Systematic Entomology doi:10.1111/syen.12275

CoL2006/BDWD - via The Interim Register of Marine and Nonmarine Genera

Pape, T.; Blagoderov, B.; Mostovski, M. B. (2011). Order Diptera Linnaeus, 1758. In Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa. 3148: 222-229. - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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