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Protorabus ✝

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Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	4764839	https://www.gbif.org/species/4764839
PaleoBioDB ID (PBDB)	153689	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=153689

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Protorabus

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
subfamily	Protorabinae	Ponomarenko 1977
genus	Protorabus	Ponomarenko 1977

Scientific Name

Protorabus Ponomarenko, 1977

Name Published In

in Arnol'di, Zherikhin, Nikritin & Ponomarenko, Trudy paleont. Inst. 161 [b]

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Protorabus	genus	belongs to Protorabinae	stated with evidence	Ponomarenko, 1977
Protorabus	genus	belongs to Carabidae	stated without evidence	Carpenter, 1992
Protorabus	genus	belongs to Protorabinae	stated with evidence	Ren, 1995

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

166 - 157 Ma
Jurassic

Last Recorded Appearance

122 - 112 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

A. G. Ponomarenko. 1977. Suborder Adephaga, Polyphaga Incertae Sedis, Infraorder Staphyliniformia, in Mezozoiskie zhestkokryiye [Mesozoic Coleoptera]. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 161:17-119

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Protorabus crassus ✝ species	accepted (GBIF) listed (PBDB)		1	133 Ma Early/Lower Cretaceous	133 Ma Early/Lower Cretaceous
Protorabus elongatus ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Protorabus kobdoensis ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Protorabus magnus ✝ species	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Protorabus minisculus ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Protorabus nigrimonticola ✝ species	accepted (GBIF) listed (PBDB)		2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Protorabus planus ✝ species	accepted (GBIF) listed (PBDB)		3	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Protorabus polyphlebius ✝ species	accepted (GBIF) listed (PBDB)		1	122 Ma Early/Lower Cretaceous	122 Ma Early/Lower Cretaceous
Protorabus tsaganensis species	accepted (GBIF)	No associated record in PBDB
Protorabus tsagensis ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous

Synonymy List

Year	Name and Author
1977	Protorabus Ponomarenko p. 72
1986	Protorabus Ponomarenko p. 91
1992	Protorabus Carpenter
1995	Protorabus Ren p. 85

References

Ponomarenko A. G. (1977) Suborder Adephaga, Polyphaga Incertae Sedis, Infraorder Staphyliniformia, in Mezozoiskie zhestkokryiye [Mesozoic Coleoptera], Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 161, 17-119

Ponomarenko A. G. (1986) Beetles. Scarabaeida (=Coleoptera), in Nasekomye v rannemelovykh ekosistemakh zapadnoy Mongolii, The Joint Soviet-Mongolian Palaeontological Expedition 28, 84-105

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655

Ren D. (1995) Insecta, Faunae and stratigraphy of Jurassic-Cretaceous in Beijing and the adjacent areas, 47-121

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

NZ cross ref or inferred from publ. title - via The Interim Register of Marine and Nonmarine Genera

Nomenclator Zoologicus (authority adjusted) - via The Interim Register of Marine and Nonmarine Genera

Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

original work (title/other) - via The Interim Register of Marine and Nonmarine Genera

in Arnol'di, Zherikhin, Nikritin & Ponomarenko, Trudy paleont. Inst. 161 [b] - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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