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Protocyrtus ✝

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Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	4786549	https://www.gbif.org/species/4786549
PaleoBioDB ID (PBDB)	139480	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=139480

Rank

genus

Taxonomy (GBIF)

Life : Animalia : Arthropoda : Insecta : Diptera : Protocyrtus

Taxonomy (PBDB)

Life : Animalia : Arthropoda : Insecta : Hymenoptera : Heloridae : Protocyrtus

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
order	Hymenoptera	Linnaeus 1758
-	Unicalcarida	Schulmeister et al. 2002
suborder	Vespina
suborder	Apocrita	Gerstaecker 1867
infraorder	Proctotrupomorpha
superfamily	Proctotrupoidea	Latreille 1802
family	Heloridae	Forster 1856
genus	Protocyrtus	Rohdendorf 1938

Scientific Name

Protocyrtus Rohdendorf, 1938

Name Published In

Trav. Inst. paléont. Acad. Sci. U. R. S. S., 7 (3)

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Protocyrtus	genus	belongs to Mesohelorinae	stated with evidence	Rasnitsyn, 1990
Protocyrtus	genus	belongs to Brachycera	stated without evidence	Carpenter, 1992
Protocyrtus	genus	belongs to Protohelorinae	stated with evidence	Johnson, 1992
Protocyrtus	genus	belongs to Mesohelorinae	stated with evidence	Zhang and Zhang, 2001
Protocyrtus	genus	belongs to Heloridae	stated with evidence	Li et al., 2017

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

166 - 157 Ma
Jurassic

Last Recorded Appearance

125 - 113 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

B. B. Rohdendorf. 1938. Dvukrylye basekomye Mezozoya Kara-Tau. I. Brachycera i chast' Nematocera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7:29-67

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Gurvanhelorus mongolicus species	accepted (GBIF)	No associated record in PBDB
Protocyrtus jurassicus ✝ species	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Protocyrtus mongolicus ✝ species	listed (PBDB)		2	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Protocyrtus parilis ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Protocyrtus turgensis ✝ species	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous

Synonymy List

Year	Name and Author
1938	Protocyrtus Rohdendorf p. 39
1986	Gurvanhelorus Rasnitsyn p. 161
1990	Protocyrtus Rasnitsyn p. 197
1992	Protocyrtus Carpenter
1992	Gurvanhelorus Johnson p. 267
1992	Protocyrtus Johnson p. 267
2001	Protocyrtus Zhang and Zhang p. 21
2013	Gurvanhelorus Shi et al. p. 138
2017	Protocyrtus Li et al. p. 481

References

Rohdendorf B. B. (1938) Dvukrylye basekomye Mezozoya Kara-Tau. I. Brachycera i chast' Nematocera, Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7, 29-67

Rasnitsyn A. P. (1986) Wasps. Vespida (=Hymenoptera), in Nasekomye v rannemelovykh ekosistemakh zapadnoy Mongolii, The Joint Soviet-Mongolian Palaeontological Expedition 28, 154-164

Rasnitsyn A. P. (1990) Hymenopterans, Vespida, in Pozdne-Mezozoyskie Nasekomye Vostochnogo Zabaykal'ya, Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 239, 177-205

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655

Johnson N. F. (1992) Catalog of world species of Proctotrupoidea, exclusive of Platygastridae (Hymenoptera), Memoirs of the American Entomological Institute 51, 1-825

Zhang H. C., Zhang J. F. (2001) Proctotrupoid wasps (Insecta, Hymenoptera) from the Yixian Formation of western Liaoning Province, Acta Micropalaeontologica Sinica 18, 11-28

Shi X. Q., Shih C. K., et al (2013) Systematic palaeontology, in New fossil helorid wasps (Insecta, Hymenoptera, Proctotrupoidea) from the Jehol Biota, China, Cretaceous Research 41, 136-142

Li L. F., Shih C. K., et al (2017) New fossil helorid wasps (Hymenoptera, Proctotrupoidea) from the Early Cretaceous of China, Alcheringa 41, 474-486

Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

as per family - via The Interim Register of Marine and Nonmarine Genera

Trav. Inst. paléont. Acad. Sci. U. R. S. S., 7 (3) - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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