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Dytiscidae
| Description | The Dytiscidae – based on the Greek dytikos (δυτικός), "able to dive" – are the predaceous diving beetles, a family of water beetles. They occur in virtually any freshwater habitat around the world, but a few species live among leaf litter. The adults of most are between 1 and 2.5 cm (0.4–1.0 in) long, though much variation is seen between species. The European Dytiscus latissimus and Brazilian Megadytes ducalis are the largest, reaching up to 4.5 cm (1.8 in) and 4.75 cm (1.9 in) respectively. In contrast, the smallest is likely the Australian Limbodessus atypicali of subterranean waters, which only is about 0.9 mm (0.035 in) long. Most are dark brown, blackish, or dark olive in color with golden highlights in some subfamilies. The larvae are commonly known as water tigers due to their voracious appetite. They have short, but sharp mandibles and immediately upon biting, they deliver digestive enzymes into prey to suck their liquefied remains. The family includes more than 4,000 described species in numerous genera. From Wikipedia article at https://en.wikipedia.org/wiki/Dytiscidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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| Source Data |
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| Rank | family | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Taxonomy (GBIF,PBDB) | Life : Animalia : Arthropoda : Insecta : Coleoptera : Dytiscidae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Taxonomic Status (GBIF) | accepted | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Classification (PBDB,GBIF) |
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| Common Name | predaceous diving beetle | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Scientific Name | Dytiscidae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Name Published In | Zool. Miscell., 3: 69 | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Opinions (PBDB) |
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| Status (PBDB) | extant | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Taxon Size (PBDB) | 225 | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Extant Size (PBDB) | 99 (44%) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| First Recorded Appearance | 183 - 174 Ma Jurassic | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Environment | terrestrial (based on Insecta) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Motility | actively mobile (based on Arthropoda) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Taphonomy | chitin (based on Insecta) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Primary Reference (PBDB) | W. E. Leach. 1815. Entomology. The Edinburgh Encyclopaedia 9:57-172 | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Common Name(s) | Waterroofkevers, Zwemtorren, Zwemkevers, Diving Beetles, Dykare, Vannkalver, Vasskalvar, Waterroofkevers, ゲンゴロウ, Predaceous Diving Beetles, Ditiques, Predacious Diving Beetles, Predaceous Diving Beetle | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Wikipedia | https://en.wikipedia.org/wiki/Dytiscidae |
External Images
Fossil Distribution
Subtaxa
| Name | Status | Common Name(s) | Fossil Occurrences | Oldest | Youngest |
|---|---|---|---|---|---|
| Agabinae subfamily | listed (PBDB) | 100 | 38.0 Ma Eocene | 0 Ma Extant | |
| Aubehydrinae subfamily | listed (PBDB) | 0 Ma Extant | |||
| Colymbetinae subfamily | listed (PBDB) | 21 | 16.0 Ma Miocene | 0 Ma Extant | |
| Copelatinae subfamily | listed (PBDB) | 9 | 38.0 Ma Eocene | 0 Ma Extant | |
| Dytiscinae subfamily | listed (PBDB) | 39 | 99.6 Ma Late/Upper Cretaceous | 0 Ma Extant | |
| Hydroporinae subfamily | listed (PBDB) | 116 | 38.0 Ma Eocene | 0 Ma Extant | |
| Laccophilinae subfamily | listed (PBDB) | 4 | 50.3 Ma Eocene | 0 Ma Extant | |
| Liadytiscinae ✝ subfamily | listed (PBDB) | 11 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Acilius genus | accepted (GBIF) listed (PBDB) | 3 | 37.2 Ma Eocene | 0 Ma Extant | |
| Aethionectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Africodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Africophilus genus | accepted (GBIF) | No associated record in PBDB | |||
| Agabetes genus | accepted (GBIF) | No associated record in PBDB | |||
| Agabinus genus | accepted (GBIF) | No associated record in PBDB | |||
| Agabus genus | accepted (GBIF) listed (PBDB) | 67 | 37.2 Ma Eocene | 0 Ma Extant | |
| Agametrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Agaporomorphus genus | accepted (GBIF) | No associated record in PBDB | |||
| Aglymbus genus | accepted (GBIF) | No associated record in PBDB | |||
| Agnoshydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Allodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Allomatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Allopachria genus | accepted (GBIF) | No associated record in PBDB | |||
| Amarodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Andex genus | accepted (GBIF) | No associated record in PBDB | |||
| Andonectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Angaragabus ✝ genus | accepted (GBIF) listed (PBDB) | 2 | 183 Ma Early Jurassic | 183 Ma Early Jurassic | |
| Anginopachria genus | accepted (GBIF) | No associated record in PBDB | |||
| Anisomeria genus | accepted (GBIF) | No associated record in PBDB | |||
| Anodocheilus genus | accepted (GBIF) | No associated record in PBDB | |||
| Antiporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Australphilus genus | accepted (GBIF) | No associated record in PBDB | |||
| Austrodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Barretthydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Batrachomatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Bidessodes genus | accepted (GBIF) | No associated record in PBDB | |||
| Bidessonotus genus | accepted (GBIF) | No associated record in PBDB | |||
| Bidessus genus | accepted (GBIF) listed (PBDB) | 1 | 37.2 Ma Eocene | 0 Ma Extant | |
| Boreonectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Borneodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Brachyvatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Bunites genus | accepted (GBIF) | No associated record in PBDB | |||
| Calicovatellus ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 20.4 Ma Miocene | 20.4 Ma Miocene | |
| Canthyporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Carabdytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Carabhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Celina genus | accepted (GBIF) | No associated record in PBDB | |||
| Chostonectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Clypeodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Coelambus genus | accepted (GBIF) | No associated record in PBDB | |||
| Coelhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Colymbetes genus | accepted (GBIF) listed (PBDB) | 21 | 16.0 Ma Miocene | 0 Ma Extant | |
| Comaldessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Copelatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Coptotomus genus | accepted (GBIF) | No associated record in PBDB | |||
| Cretodytes ✝ genus | accepted (GBIF) listed (PBDB) | 2 | 129 Ma Early/Lower Cretaceous | 93.9 Ma Late/Upper Cretaceous | |
| Crinodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Cybister genus | accepted (GBIF) listed (PBDB) | 11 | 20.0 Ma Miocene | 0 Ma Extant | |
| Darwinhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Deronectes genus | accepted (GBIF) listed (PBDB) | 1 | 0.13 Ma Pleistocene | 0 Ma Extant | |
| Derovatellus genus | accepted (GBIF) listed (PBDB) | 0 Ma Extant | |||
| Desmopachria genus | accepted (GBIF) | No associated record in PBDB | |||
| Dimitshydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Dytiscus genus | accepted (GBIF) listed (PBDB) | 16 | 38.0 Ma Eocene | 0 Ma Extant | |
| Ereboporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Eretes genus | accepted (GBIF) listed (PBDB) | 1 | 33.9 Ma Oligocene | 0 Ma Extant | |
| Etruscodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Exocelina genus | accepted (GBIF) | No associated record in PBDB | |||
| Fontidessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Geodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Gibbidessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Glareadessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Glessaria ✝ genus | accepted (GBIF) listed (PBDB) | 2 | 38.0 Ma Eocene | 38.0 Ma Eocene | |
| Graphoderus genus | accepted (GBIF) listed (PBDB) | 4 | 16.0 Ma Miocene | 0 Ma Extant | |
| Graptodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Haideoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hemibidessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Heroceras genus | accepted (GBIF) | No associated record in PBDB | |||
| Herophydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Heterhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Heterosternuta genus | accepted (GBIF) | No associated record in PBDB | |||
| Hoperius genus | accepted (GBIF) | No associated record in PBDB | |||
| Hovahydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Huxelhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hyderodes genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydrocolus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydrodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydrodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydroglyphus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydronebrius genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydropeplus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hydroporus genus | accepted (GBIF) listed (PBDB) | 51 | 38.0 Ma Eocene | 0 Ma Extant | |
| Hydrotrupes genus | accepted (GBIF) listed (PBDB) | 1 | 38.0 Ma Eocene | 0 Ma Extant | |
| Hydrovatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hygrotus genus | accepted (GBIF) listed (PBDB) | 20 | 37.2 Ma Eocene | 0 Ma Extant | |
| Hyphoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hyphovatus genus | accepted (GBIF) | No associated record in PBDB | |||
| Hyphydrus genus | accepted (GBIF) listed (PBDB) | 0 Ma Extant | |||
| Hypodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Iberoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Ilybiosoma genus | accepted (GBIF) | No associated record in PBDB | |||
| Ilybius genus | accepted (GBIF) listed (PBDB) | 26 | 5.33 Ma Miocene | 0 Ma Extant | |
| Incomptodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Japanolaccophilus genus | accepted (GBIF) listed (PBDB) | 1 | 38.0 Ma Eocene | 0 Ma Extant | |
| Kakadudessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Kuschelydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Laccodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Laccomimus genus | accepted (GBIF) | No associated record in PBDB | |||
| Lacconectus genus | accepted (GBIF) | No associated record in PBDB | |||
| Laccophilus genus | accepted (GBIF) listed (PBDB) | 3 | 50.3 Ma Eocene | 0 Ma Extant | |
| Laccoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Laccornellus genus | accepted (GBIF) | No associated record in PBDB | |||
| Laccornis genus | accepted (GBIF) | No associated record in PBDB | |||
| Lancetes genus | accepted (GBIF) | No associated record in PBDB | |||
| Leiodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Leuronectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Liadroporus ✝ genus | accepted (GBIF) listed (PBDB) | 2 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Liadytiscus ✝ genus | accepted (GBIF) listed (PBDB) | 3 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Liadyxianus ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Limbodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Liodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Lioporeus genus | accepted (GBIF) | No associated record in PBDB | |||
| Liopterus genus | accepted (GBIF) | No associated record in PBDB | |||
| Madaglymbus genus | accepted (GBIF) | No associated record in PBDB | |||
| Matus genus | accepted (GBIF) | No associated record in PBDB | |||
| Megadytes genus | accepted (GBIF) listed (PBDB) | 1 | 0.13 Ma Pleistocene | 0 Ma Extant | |
| Megaporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Meladema genus | accepted (GBIF) | No associated record in PBDB | |||
| Melanodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Mesoderus ✝ genus | accepted (GBIF) listed (PBDB) | 4 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Mesodytes ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Metaporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Methles genus | accepted (GBIF) listed (PBDB) | 1 | 16.0 Ma Miocene | 0 Ma Extant | |
| Microdessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Microdytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Microhydrodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Miodytiscus ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 37.2 Ma Eocene | 37.2 Ma Eocene | |
| Morimotoa genus | accepted (GBIF) | No associated record in PBDB | |||
| Napodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Nebrioporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Necterosoma genus | accepted (GBIF) | No associated record in PBDB | |||
| Neobidessodes genus | accepted (GBIF) | No associated record in PBDB | |||
| Neobidessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Neoclypeodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Neoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Neoscutopterus genus | accepted (GBIF) | No associated record in PBDB | |||
| Neptosternus genus | accepted (GBIF) | No associated record in PBDB | |||
| Notaticus genus | accepted (GBIF) | No associated record in PBDB | |||
| Novadessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Onychohydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Oreodytes genus | accepted (GBIF) listed (PBDB) | 8 | 28.1 Ma Oligocene | 0 Ma Extant | |
| Pachydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Pachynectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Palaeodytes ✝ genus | accepted (GBIF) listed (PBDB) | 3 | 166 Ma Middle Jurassic | 125 Ma Early/Lower Cretaceous | |
| Palaeogyrinus ✝ genus | accepted (GBIF) listed (PBDB) | 2 | 28.4 Ma Oligocene | 28.1 Ma Oligocene | |
| Papuadessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Paroster genus | accepted (GBIF) | No associated record in PBDB | |||
| Peschetius genus | accepted (GBIF) | No associated record in PBDB | |||
| Petrodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Philaccolilus genus | accepted (GBIF) | No associated record in PBDB | |||
| Philaccolus genus | accepted (GBIF) | No associated record in PBDB | |||
| Philodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Phreatodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Platambus genus | accepted (GBIF) listed (PBDB) | 2 | 0.78 Ma Pleistocene | 0 Ma Extant | |
| Platydytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Platynectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Porhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Potamonectes genus | listed (PBDB) | 4 | 0.78 Ma Pleistocene | 0 Ma Extant | |
| Primospes genus | accepted (GBIF) | No associated record in PBDB | |||
| Procoelambus ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 28.1 Ma Oligocene | 28.1 Ma Oligocene | |
| Prodaticus genus | accepted (GBIF) | No associated record in PBDB | |||
| Pseuduvarus genus | accepted (GBIF) | No associated record in PBDB | |||
| Queda genus | accepted (GBIF) | No associated record in PBDB | |||
| Regimbartina genus | accepted (GBIF) | No associated record in PBDB | |||
| Rhantaticus genus | accepted (GBIF) | No associated record in PBDB | |||
| Rhantus genus | accepted (GBIF) listed (PBDB) | 6 | 1.81 Ma Pleistocene | 0 Ma Extant | |
| Rhithrodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Rompindessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Rugosus genus | accepted (GBIF) | No associated record in PBDB | |||
| Sandracottus genus | accepted (GBIF) | No associated record in PBDB | |||
| Sanfilippodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Scarodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Schistomerus ✝ genus | accepted (GBIF) listed (PBDB) | 21 | 20.4 Ma Miocene | 20.4 Ma Miocene | |
| Sekaliporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Senilites genus | accepted (GBIF) | No associated record in PBDB | |||
| Sharphydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Siamoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Siettitia genus | accepted (GBIF) | No associated record in PBDB | |||
| Sinodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Sinoporus ✝ genus | accepted (GBIF) listed (PBDB) | 1 | 125 Ma Early/Lower Cretaceous | 125 Ma Early/Lower Cretaceous | |
| Spanglerodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Spencerhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Sternhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Sternopriscus genus | accepted (GBIF) | No associated record in PBDB | |||
| Stictonectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Stictotarsus genus | accepted (GBIF) | No associated record in PBDB | |||
| Stygoporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Suphrodytes genus | accepted (GBIF) | No associated record in PBDB | |||
| Tepuidessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Terradessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Thermonectus genus | accepted (GBIF) | No associated record in PBDB | |||
| Tikoloshanes genus | accepted (GBIF) | No associated record in PBDB | |||
| Tiporus genus | accepted (GBIF) | No associated record in PBDB | |||
| Trichonectes genus | accepted (GBIF) | No associated record in PBDB | |||
| Trogloguignotus genus | accepted (GBIF) | No associated record in PBDB | |||
| Tyndallhydrus genus | accepted (GBIF) | No associated record in PBDB | |||
| Typhlodessus genus | accepted (GBIF) | No associated record in PBDB | |||
| Uvarus genus | accepted (GBIF) | No associated record in PBDB | |||
| Vatellus genus | accepted (GBIF) | No associated record in PBDB | |||
| Yola genus | accepted (GBIF) | No associated record in PBDB | |||
| Yolina genus | accepted (GBIF) | No associated record in PBDB | |||
| Belladessus femineus species | accepted (GBIF) | No associated record in PBDB | |||
| Belladessus puella species | accepted (GBIF) | No associated record in PBDB | |||
| Brinckoides biguttatus species | accepted (GBIF) | No associated record in PBDB | |||
| Deroncectes minax species | accepted (GBIF) | No associated record in PBDB | |||
| Platynectus buruensis species | accepted (GBIF) | No associated record in PBDB | |||
Synonymy List
| Year | Name and Author |
|---|---|
| 1856 | Necticus palustris Aymard p. 234 |
| 1861 | Dytiscidae LeConte p. 38 |
| 1981 | Dytiscidae Crowson p. 694 |
| 1992 | Dytiscidae Carpenter |
| 1995 | Dytiscidae Lawrence and Newton, Jr. |
| 2003 | Dytiscidae Wappler |
| 2011 | Dytiscidae Bouchard et al. p. 162 |
| 2019 | Dytiscidae Yang et al. p. 2 |
References
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
