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Arctocyonidae ✝

Description

Arctocyonidae (from Greek arktos kyôn, "bear/dog-like") has been defined as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene (66 to 50 million years ago).Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)3239418https://www.gbif.org/species/3239418
PaleoBioDB ID (PBDB)42198https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=42198
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Chordata : Mammalia : Arctocyonidae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-TriploblasticaLankester 1877
-NephrozoaJondelius et al. 2002
-DeuterostomiaGrobben 1908
phylumChordataHaeckel 1847
subphylumVertebrataLamarck 1801
superclassGnathostomataGegenbauer 1874
-Osteichthyes
-SarcopterygiiRomer 1955
subclassDipnotetrapodomorpha
subclassTetrapodomorpha
-Tetrapoda
-ReptiliomorphaSäve-Söderbergh 1934
-AnthracosauriaSäve-Söderbergh 1934
-Batrachosauria
-Cotylosauria
-AmniotaHaeckel 1866
-SynapsidaOsborn 1903
-Therapsida
infraorderCynodontia
-EpicynodontiaHopson and Kitching 2001
infraorderEucynodontiaKemp 1982
-ProbainognathiaHopson 1990
-MammaliamorphaRowe 1988
-MammaliaformesRowe 1988
classMammaliaLinnaeus 1758
subclassTribosphenida
infraclassEutheria
familyArctocyonidaeMurray 1866
Common Namecondylarth
Scientific NameArctocyonidae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Arctocyonidaefamilybelongs to Creodontastated with evidenceCope, 1880
Arctocyonidaefamilybelongs to Creodontastated without evidenceCope, 1880
Arctocyonidaefamilybelongs to Arctocyonoideastated without evidenceSavage, 1965
Arctocyonidaefamilybelongs to Condylarthrastated without evidenceSloan and Van Valen, 1965
Arctocyonidaefamilybelongs to Creodontastated without evidenceGazin, 1969
Arctocyonidaefamilybelongs to Arctocyonoideastated without evidenceKielan-Jaworowska et al., 1979
Arctocyonidaefamilybelongs to CondylarthraimpliedCarroll, 1988
Arctocyonidaefamilybelongs to Ungulatastated with evidenceArchibald, 1998
Arctocyonidaefamilybelongs to Arctocyoniastated without evidenceThewissen et al., 2001
Arctocyonidaefamilybelongs to Condylarthrastated without evidenceHooker and Millbank, 2001
Arctocyonidaefamilybelongs to Procreodistated without evidenceMcKenna and Lofgren, 2003
Arctocyonidaefamilybelongs to Condylarthrastated without evidenceKielan-Jaworowska et al., 2004
Arctocyonidaefamilybelongs to Eutheriastated with evidenceTabuce et al., 2011
Arctocyonidaefamilybelongs to Condylarthrastated without evidenceRose et al., 2012
Status (PBDB)extinct
Taxon Size (PBDB)116
First Recorded Appearance72.1 - 66.0 Ma
Late/Upper Cretaceous
Last Recorded Appearance48.6 - 37.2 Ma
Eocene
Environmentterrestrial (based on Eutheria)
Motilityactively mobile (based on Osteichthyes)
Dietomnivore
Ecospace Commentsominvorous (Wilson 1975, p. 368); ground dwelling life habit is based on Arctocyon, but Chriacus was scansorial (Archibald 1998)
Taphonomyphosphatic (based on Vertebrata)
Primary Reference (PBDB)R. L. Carroll. 1988. Vertebrate Paleontology and Evolution
Common Name(s) Condylarth
Wikipediahttps://en.wikipedia.org/wiki/Arctocyonidae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Arctocyoninae
subfamily
listed (PBDB)
Loxolophinae
subfamily
listed (PBDB)
Anacodon
genus
accepted (GBIF)
listed (PBDB)
2961.7 Ma
Paleocene
55.8 Ma
Eocene
Aphanocyon
genus
accepted (GBIF)
listed (PBDB)
461.7 Ma
Paleocene
58.7 Ma
Paleocene
Arctocyon
genus
accepted (GBIF)
listed (PBDB)
7863.3 Ma
Paleocene
56.8 Ma
Paleocene
Arctocyonides
genus
accepted (GBIF)
listed (PBDB)
761.6 Ma
Paleocene
56.0 Ma
Paleocene
Baioconodon
genus
accepted (GBIF)
listed (PBDB)
5272.1 Ma
Late/Upper Cretaceous
66.0 Ma
Late/Upper Cretaceous
Carcinodon
genus
accepted (GBIF)
listed (PBDB)
666.0 Ma
Late/Upper Cretaceous
66.0 Ma
Late/Upper Cretaceous
Chriacus
genus
accepted (GBIF)
listed (PBDB)
16466.0 Ma
Late/Upper Cretaceous
55.8 Ma
Eocene
Claenodon
genus
accepted (GBIF)
listed (PBDB)
1363.3 Ma
Paleocene
61.7 Ma
Paleocene
Colpoclaenus
genus
accepted (GBIF)
listed (PBDB)
1763.3 Ma
Paleocene
61.7 Ma
Paleocene
Desmatoclaenus
genus
accepted (GBIF)
listed (PBDB)
1566.0 Ma
Late/Upper Cretaceous
61.7 Ma
Paleocene
Deuterogonodon
genus
accepted (GBIF)
listed (PBDB)
663.3 Ma
Paleocene
63.3 Ma
Paleocene
Heteroborus
genus
accepted (GBIF)
listed (PBDB)
Hyodectes
genus
accepted (GBIF)
listed (PBDB)
Karakia
genus
accepted (GBIF)
listed (PBDB)
155.8 Ma
Eocene
55.8 Ma
Eocene
Lambertocyon
genus
accepted (GBIF)
listed (PBDB)
2161.7 Ma
Paleocene
56.8 Ma
Paleocene
Landenodon
genus
accepted (GBIF)
listed (PBDB)
1259.2 Ma
Paleocene
55.8 Ma
Eocene
Loxolophus
genus
accepted (GBIF)
listed (PBDB)
4566.0 Ma
Late/Upper Cretaceous
63.3 Ma
Paleocene
Mentoclaenodon
genus
accepted (GBIF)
listed (PBDB)
461.7 Ma
Paleocene
61.6 Ma
Paleocene
Mimotricentes
genus
accepted (GBIF)
listed (PBDB)
1763.3 Ma
Paleocene
61.7 Ma
Paleocene
Oxyprimus
genus
accepted (GBIF)
listed (PBDB)
3866.0 Ma
Late/Upper Cretaceous
61.7 Ma
Paleocene
Oxytomodon
genus
accepted (GBIF)
listed (PBDB)
163.3 Ma
Paleocene
63.3 Ma
Paleocene
Paratriisodon
genus
accepted (GBIF)
listed (PBDB)
248.6 Ma
Eocene
48.6 Ma
Eocene
Princetonia
genus
accepted (GBIF)
listed (PBDB)
1761.7 Ma
Paleocene
55.8 Ma
Eocene
Prothryptacodon
genus
accepted (GBIF)
listed (PBDB)
1466.0 Ma
Late/Upper Cretaceous
61.7 Ma
Paleocene
Protungulatum
genus
accepted (GBIF)
listed (PBDB)
6270.6 Ma
Late/Upper Cretaceous
66.0 Ma
Late/Upper Cretaceous
Sigynorum
genus
accepted (GBIF)
listed (PBDB)
266.0 Ma
Late/Upper Cretaceous
66.0 Ma
Late/Upper Cretaceous
Thryptacodon
genus
accepted (GBIF)
listed (PBDB)
16763.3 Ma
Paleocene
50.3 Ma
Eocene

Synonymy List

YearName and Author
1866Arctocyonidae Murray
1880Arctocyonidae Cope
1889Arctocyonidae Cope p. 876
1891Arctocyonidae Flower and Lydekker p. 91
1902Arctocyonidae Hay p. 746
1965Arctocyonidae Savage p. 243
1965Arctocyonidae Sloan and Van Valen p. 226
1969Arctocyonidae Gazin p. 7
1979Arctocyonidae Kielan-Jaworowska et al. p. 223
YearName and Author
1988Arctocyonidae Carroll
1998Arctocyonidae Archibald p. 299
2001Arctocyonidae Hooker and Millbank
2001Arctocyonidae Thewissen et al. p. 350 figs. Table 1
2002Arctocyonidae Scott et al. p. 695
2003Arctocyonidae McKenna and Lofgren p. 635
2004Arctocyonidae Kielan-Jaworowska et al. pp. 495, 513-514
2011Arctocyonidae Tabuce et al. p. 152
2012Arctocyonidae Rose et al.

References

Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698
Cope E. D. On the genera of the Creodonta, Proceedings of the American Philosophical Society 19, 76-82
Cope E. D. Synopsis of the families of Vertebrata, The American Naturalist 23, 1-29
Hay O. P. (1902) , Bibliography and Catalogue of the Fossil Vertebrata of North America. Bulletin of the United States Geological Survey 179, 1-868
Savage R. J. G. (1965) Fossil Mammals of Africa: 19 The Miocene Carnivora of East Africa, Bulletin of the British Museum (Natural History) Geology 10 8, 241-316
Sloan R. E., Van Valen L. (1965) Cretaceous mammals from Montana, Science 148 3667, 220-227
Gazin C. L. (1969) A new occurrence of Paleocene mammals in the Evanston Formation, southwestern Wyoming, Smithsonian Contributions to Paleobiology 2, 1-16
Kielan-Jaworowska Z., Bown T. M., et al (1979) Eutheria, Mesozoic mammals: the first two-thirds of mammalian history, 221-258
Archibald J. D. (1998) Archaic ungulates ("Condylarthra"), Evolution of Tertiary Mammals of North America 1, 292-331
Hooker J. J., Millbank C. (2001) A Cernaysian mammal from the Upnor Formation (Late Palaeocene, Herne Bay, UK) and its implications for correlation, Proceedings of the Geologists' Association 112, 331-338
Thewissen J. G. M., Williams E. M., et al (2001) Eocene mammal faunas from northern Indo-Pakistan, Journal of Vertebrate Paleontology 21 2, 347-366
Scott C. S., Fox R. C., et al (2002) New earliest Tiffanian (late Paleocene) mammals from Cochrane 2, southwestern Alberta, Canada, Acta Palaeontologica Polonica 47 4, 691-704
McKenna M. C., Lofgren D. L. (2003) Mimotricentes tedfordi, a new arctocyonid from the late Paleocene of California, Bulletin of the American Museum of Natural History 279, 632-643
Kielan-Jaworowska Z., Cifelli R. L., et al (2004) Mammals from the age of dinosaurs: Origins, evolution, and structure, Columbia University Press, New York, 1-630
Tabuce R., Clavel J., et al (2011) A structural intermediate between triisodontids and mesonychians (Mammalia, Acreodi) from the earliest Eocene of Portugal, Naturwissenschaften 98, 145-155
Rose K. D., Chew A. E., et al (2012) Earliest Eocene mammalian fauna from the Paleocene-Eocene Thermal Maximum at Sand Creek Divide, southern Bighorn Basin, Wyoming, University of Michigan Papers on Paleontology 36, 1-122
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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