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Helophorus

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Description

Helophorus ís the only genus in the beetle family Helophoridae (traditionally included within Hydrophilidae as the subfamily Helophorinae). They are small insects, found mainly in the Holarctic region (150 occur in Palearctic and 41 species in North America), but two or three species also live in the Afrotropical region, Central America and one in the Indomalayan region (northern India).

From Wikipedia article at https://en.wikipedia.org/wiki/Helophorus, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3263442	https://www.gbif.org/species/3263442
PaleoBioDB ID (PBDB)	193933	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=193933

Rank

genus

Taxonomy (GBIF)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : Helophorus

Taxonomy (PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : Helophorus

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Polyphaga	Emery 1886
infraorder	Staphyliniformia	Lameere 1900
superfamily	Hydrophiloidea	Latreille 1802
family	Helophoridae
genus	Helophorus	Fabricius 1775

Scientific Name

Helophorus Fabricius, 1775

Name Published In

Fabricius, Johann C. 1775. Systema Entomologiae: sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Officina Libraria Kortii, Flensburgi et Lipsiae.: i–xvi; 1–832.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Helophorus	genus	belongs to Helophorina	stated with evidence	Sahlberg, 1880
Helophorus	genus	belongs to Hydrophilidae	stated with evidence	Scudder, 1900
Helophorus	genus	belongs to Helophorinae	stated with evidence	Bouchard et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

27 (75%)

First Recorded Appearance

152 - 145 Ma
Late Jurassic

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

J. C. Fabricius. 1775. Systema Entomologiae, Sistens Insectorum Classes, Ordines, Genera, Species Adiectis Synonymis, Locis, Descriptionibus, Observationibus

Wikipedia

https://en.wikipedia.org/wiki/Helophorus

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Helophorus (Atracthelophorus) subgenus	listed (PBDB)		2	0.13 Ma Pleistocene	0 Ma Extant
Helophorus (Gephelophorus) subgenus	listed (PBDB)		5	23.0 Ma Miocene	0 Ma Extant
Helophorus (Meghelophorus) subgenus	listed (PBDB)		8	2.59 Ma Pleistocene	0 Ma Extant
Helophorus (Mesohelophorus) ✝ subgenus	listed (PBDB)		2	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Helophorus (Mesosperchus) ✝ subgenus	listed (PBDB)		17	152 Ma Late Jurassic	125 Ma Early/Lower Cretaceous
Helophorus (Orphelophorus) subgenus	listed (PBDB)		2	0.78 Ma Pleistocene	0 Ma Extant
Helophorus (Rhopalhelophorus) subgenus	listed (PBDB)				0 Ma Extant
Helophorus (Trichelophorus) subgenus	listed (PBDB)				0 Ma Extant
Helophorus abeillei species	accepted (GBIF)	No associated record in PBDB
Helophorus aequalis species	accepted (GBIF)	No associated record in PBDB
Helophorus aethiopicus species	accepted (GBIF)	No associated record in PBDB
Helophorus aethiops species	accepted (GBIF)	No associated record in PBDB
Helophorus algiricus species	accepted (GBIF)	No associated record in PBDB
Helophorus altaicus species	accepted (GBIF)	No associated record in PBDB
Helophorus alternans species	accepted (GBIF)	No associated record in PBDB
Helophorus alternatus species	accepted (GBIF)	No associated record in PBDB
Helophorus angusi species	accepted (GBIF)	No associated record in PBDB
Helophorus angustatus species	accepted (GBIF)	No associated record in PBDB
Helophorus angusticollis species	accepted (GBIF)	No associated record in PBDB
Helophorus apfelbecki species	accepted (GBIF)	No associated record in PBDB
Helophorus aquaticus species	accepted (GBIF)	No associated record in PBDB
Helophorus aquila species	accepted (GBIF)	No associated record in PBDB
Helophorus arcticus species	accepted (GBIF)	No associated record in PBDB
Helophorus armeniacus species	accepted (GBIF)	No associated record in PBDB
Helophorus artus species	accepted (GBIF)	No associated record in PBDB
Helophorus arvernicus species	accepted (GBIF)	No associated record in PBDB
Helophorus asperatus species	accepted (GBIF)	No associated record in PBDB
Helophorus aspericollis species	accepted (GBIF)	No associated record in PBDB
Helophorus asturiensis species	accepted (GBIF)	No associated record in PBDB
Helophorus atlantis species	accepted (GBIF)	No associated record in PBDB
Helophorus auricollis species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus auriculatus species	accepted (GBIF)	No associated record in PBDB
Helophorus bameuli species	accepted (GBIF)	No associated record in PBDB
Helophorus banghaasi species	accepted (GBIF)	No associated record in PBDB
Helophorus barbarae species	accepted (GBIF)	No associated record in PBDB
Helophorus beibienkoi species	accepted (GBIF)	No associated record in PBDB
Helophorus bergrothi species	accepted (GBIF)	No associated record in PBDB
Helophorus biltoni species	accepted (GBIF)	No associated record in PBDB
Helophorus brevipalpis species	accepted (GBIF) listed (PBDB)		3	0.13 Ma Pleistocene	0 Ma Extant
Helophorus brevitarsis species	accepted (GBIF)	No associated record in PBDB
Helophorus browni species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus californicus species	accepted (GBIF)	No associated record in PBDB
Helophorus calpensis species	accepted (GBIF)	No associated record in PBDB
Helophorus carsoni species	accepted (GBIF)	No associated record in PBDB
Helophorus chamberlaini species	accepted (GBIF)	No associated record in PBDB
Helophorus cincticollis species	accepted (GBIF)	No associated record in PBDB
Helophorus columbianus species	accepted (GBIF)	No associated record in PBDB
Helophorus confrater species	accepted (GBIF)	No associated record in PBDB
Helophorus coopei species	accepted (GBIF)	No associated record in PBDB
Helophorus costalis species	accepted (GBIF)	No associated record in PBDB
Helophorus costulatus species	accepted (GBIF)	No associated record in PBDB
Helophorus crinitus species	accepted (GBIF)	No associated record in PBDB
Helophorus croaticus species	accepted (GBIF)	No associated record in PBDB
Helophorus cuspifer species	accepted (GBIF)	No associated record in PBDB
Helophorus daedalus species	accepted (GBIF)	No associated record in PBDB
Helophorus dalmatinus species	accepted (GBIF)	No associated record in PBDB
Helophorus difficilis species	accepted (GBIF)	No associated record in PBDB
Helophorus discrepans species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus discretus species	accepted (GBIF)	No associated record in PBDB
Helophorus dixoni species	accepted (GBIF)	No associated record in PBDB
Helophorus dorsalis species	accepted (GBIF)	No associated record in PBDB
Helophorus dracomontanus species	accepted (GBIF)	No associated record in PBDB
Helophorus eclectus species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus elizabethae species	accepted (GBIF)	No associated record in PBDB
Helophorus erzurumicus species	accepted (GBIF)	No associated record in PBDB
Helophorus faustianus species	accepted (GBIF)	No associated record in PBDB
Helophorus fauveli species	accepted (GBIF)	No associated record in PBDB
Helophorus fenderi species	accepted (GBIF)	No associated record in PBDB
Helophorus flavipes species	accepted (GBIF)	No associated record in PBDB
Helophorus fortis species	accepted (GBIF)	No associated record in PBDB
Helophorus fosteri species	accepted (GBIF)	No associated record in PBDB
Helophorus frater species	accepted (GBIF)	No associated record in PBDB
Helophorus frosti species	accepted (GBIF)	No associated record in PBDB
Helophorus fulgidicollis species	accepted (GBIF)	No associated record in PBDB
Helophorus furius species	accepted (GBIF)	No associated record in PBDB
Helophorus glacialis species	accepted (GBIF)	No associated record in PBDB
Helophorus gracilis species	accepted (GBIF)	No associated record in PBDB
Helophorus grandis species	accepted (GBIF)	No associated record in PBDB
Helophorus granularis species	accepted (GBIF)	No associated record in PBDB
Helophorus gratus species	accepted (GBIF)	No associated record in PBDB
Helophorus griseus species	accepted (GBIF)	No associated record in PBDB
Helophorus gurjevae species	accepted (GBIF)	No associated record in PBDB
Helophorus guttulus species	accepted (GBIF)	No associated record in PBDB
Helophorus hammondi species	accepted (GBIF)	No associated record in PBDB
Helophorus hatchi species	accepted (GBIF)	No associated record in PBDB
Helophorus helenae species	accepted (GBIF)	No associated record in PBDB
Helophorus hilaris species	accepted (GBIF)	No associated record in PBDB
Helophorus hirsutiventris species	accepted (GBIF)	No associated record in PBDB
Helophorus hispanicus species	accepted (GBIF)	No associated record in PBDB
Helophorus illustris species	accepted (GBIF)	No associated record in PBDB
Helophorus imaensis species	accepted (GBIF)	No associated record in PBDB
Helophorus inceptivus species	accepted (GBIF)	No associated record in PBDB
Helophorus inexpectatus species	accepted (GBIF)	No associated record in PBDB
Helophorus inflectus species	accepted (GBIF)	No associated record in PBDB
Helophorus jaechi species	accepted (GBIF)	No associated record in PBDB
Helophorus jocoteroi species	accepted (GBIF)	No associated record in PBDB
Helophorus karatavicus species	accepted (GBIF)	No associated record in PBDB
Helophorus kaszabianus species	accepted (GBIF)	No associated record in PBDB
Helophorus kayae species	accepted (GBIF)	No associated record in PBDB
Helophorus kazakhstanicus species	accepted (GBIF)	No associated record in PBDB
Helophorus kerimi species	accepted (GBIF)	No associated record in PBDB
Helophorus kervillei species	accepted (GBIF)	No associated record in PBDB
Helophorus khnzoriani species	accepted (GBIF)	No associated record in PBDB
Helophorus kirgisicus species	accepted (GBIF)	No associated record in PBDB
Helophorus korotyaevi species	accepted (GBIF)	No associated record in PBDB
Helophorus kozlovi species	accepted (GBIF)	No associated record in PBDB
Helophorus kryzhanovskii species	accepted (GBIF)	No associated record in PBDB
Helophorus lacustris species	accepted (GBIF) listed (PBDB)		5	0.13 Ma Pleistocene	0 Ma Extant
Helophorus laferi species	accepted (GBIF)	No associated record in PBDB
Helophorus lamicola species	accepted (GBIF)	No associated record in PBDB
Helophorus lapponicus species	accepted (GBIF)	No associated record in PBDB
Helophorus laticollis species	accepted (GBIF)	No associated record in PBDB
Helophorus latipenis species	accepted (GBIF)	No associated record in PBDB
Helophorus lecontei species	accepted (GBIF)	No associated record in PBDB
Helophorus ledatus species	accepted (GBIF)	No associated record in PBDB
Helophorus leechi species	accepted (GBIF)	No associated record in PBDB
Helophorus leontis species	accepted (GBIF)	No associated record in PBDB
Helophorus lewisi species	accepted (GBIF)	No associated record in PBDB
Helophorus liguricus species	accepted (GBIF)	No associated record in PBDB
Helophorus limbatus species	accepted (GBIF)	No associated record in PBDB
Helophorus linearis species	accepted (GBIF)	No associated record in PBDB
Helophorus linearoides species	accepted (GBIF)	No associated record in PBDB
Helophorus lineatus species	accepted (GBIF)	No associated record in PBDB
Helophorus longipennis species	accepted (GBIF)	No associated record in PBDB
Helophorus longitarsis species	accepted (GBIF)	No associated record in PBDB
Helophorus maculatus species	accepted (GBIF)	No associated record in PBDB
Helophorus manchuricus species	accepted (GBIF)	No associated record in PBDB
Helophorus marginicollis species	accepted (GBIF)	No associated record in PBDB
Helophorus maritimus species	accepted (GBIF)	No associated record in PBDB
Helophorus matsumurai species	accepted (GBIF)	No associated record in PBDB
Helophorus meighenensis species	accepted (GBIF)	No associated record in PBDB
Helophorus mervensis species	accepted (GBIF)	No associated record in PBDB
Helophorus micans species	accepted (GBIF)	No associated record in PBDB
Helophorus milleri species	accepted (GBIF)	No associated record in PBDB
Helophorus minutus species	accepted (GBIF)	No associated record in PBDB
Helophorus mongoliensis species	accepted (GBIF)	No associated record in PBDB
Helophorus montanus species	accepted (GBIF)	No associated record in PBDB
Helophorus montenegrinus species	accepted (GBIF)	No associated record in PBDB
Helophorus nanus species	accepted (GBIF)	No associated record in PBDB
Helophorus nevadensis species	accepted (GBIF)	No associated record in PBDB
Helophorus niger species	accepted (GBIF)	No associated record in PBDB
Helophorus nigricans species	accepted (GBIF)	No associated record in PBDB
Helophorus nitiduloides species	accepted (GBIF)	No associated record in PBDB
Helophorus nitidulus species	accepted (GBIF)	No associated record in PBDB
Helophorus nivalis species	accepted (GBIF)	No associated record in PBDB
Helophorus nivicola species	accepted (GBIF)	No associated record in PBDB
Helophorus nubilus species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus oblongus species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus obscurellus species	accepted (GBIF)	No associated record in PBDB
Helophorus obscurus species	accepted (GBIF)	No associated record in PBDB
Helophorus occidentalis species	accepted (GBIF)	No associated record in PBDB
Helophorus orchymonti species	accepted (GBIF)	No associated record in PBDB
Helophorus oregonus species	accepted (GBIF)	No associated record in PBDB
Helophorus orientalis species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Helophorus oscillator species	accepted (GBIF)	No associated record in PBDB
Helophorus oxygonus species	accepted (GBIF)	No associated record in PBDB
Helophorus palaeosibiricus species	accepted (GBIF)	No associated record in PBDB
Helophorus pallidipennis species	accepted (GBIF)	No associated record in PBDB
Helophorus pallidus species	accepted (GBIF)	No associated record in PBDB
Helophorus parajacutus species	accepted (GBIF)	No associated record in PBDB
Helophorus paraminutus species	accepted (GBIF)	No associated record in PBDB
Helophorus paramontanus species	accepted (GBIF)	No associated record in PBDB
Helophorus parasplendidus species	accepted (GBIF)	No associated record in PBDB
Helophorus peyerimhoffi species	accepted (GBIF)	No associated record in PBDB
Helophorus pici species	accepted (GBIF)	No associated record in PBDB
Helophorus pictus species	accepted (GBIF)	No associated record in PBDB
Helophorus pitcheri species	accepted (GBIF)	No associated record in PBDB
Helophorus ponticus species	accepted (GBIF)	No associated record in PBDB
Helophorus poppii species	accepted (GBIF)	No associated record in PBDB
Helophorus porculus species	accepted (GBIF)	No associated record in PBDB
Helophorus praenanus species	accepted (GBIF) listed (PBDB)		3	0.13 Ma Pleistocene	0 Ma Extant
Helophorus pumilio species	accepted (GBIF)	No associated record in PBDB
Helophorus puncticollis species	accepted (GBIF)	No associated record in PBDB
Helophorus redtenbacheri species	accepted (GBIF)	No associated record in PBDB
Helophorus regularis species	accepted (GBIF)	No associated record in PBDB
Helophorus richterae species	accepted (GBIF)	No associated record in PBDB
Helophorus rigescens ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Helophorus rinki species	accepted (GBIF)	No associated record in PBDB
Helophorus robertsi species	accepted (GBIF)	No associated record in PBDB
Helophorus rufipes species	accepted (GBIF)	No associated record in PBDB
Helophorus salinus species	accepted (GBIF)	No associated record in PBDB
Helophorus schmidti species	accepted (GBIF)	No associated record in PBDB
Helophorus schoedli species	accepted (GBIF)	No associated record in PBDB
Helophorus schuhi species	accepted (GBIF)	No associated record in PBDB
Helophorus seidlitzii species	accepted (GBIF)	No associated record in PBDB
Helophorus sempervarians species	accepted (GBIF) listed (PBDB)		8	0.13 Ma Pleistocene	0 Ma Extant
Helophorus ser species	accepted (GBIF)	No associated record in PBDB
Helophorus shatrovskyi species	accepted (GBIF)	No associated record in PBDB
Helophorus sibiricus species	accepted (GBIF)	No associated record in PBDB
Helophorus similis species	accepted (GBIF)	No associated record in PBDB
Helophorus singularis species	accepted (GBIF)	No associated record in PBDB
Helophorus sinoglacialis species	accepted (GBIF)	No associated record in PBDB
Helophorus smetanai species	accepted (GBIF)	No associated record in PBDB
Helophorus splendidus species	accepted (GBIF) listed (PBDB)		9	2.59 Ma Pleistocene	0 Ma Extant
Helophorus strandi species	accepted (GBIF)	No associated record in PBDB
Helophorus strigifrons species	accepted (GBIF)	No associated record in PBDB
Helophorus subarcuatus species	accepted (GBIF)	No associated record in PBDB
Helophorus subcarinatus species	accepted (GBIF)	No associated record in PBDB
Helophorus syriacus species	accepted (GBIF)	No associated record in PBDB
Helophorus tarsalis species	accepted (GBIF)	No associated record in PBDB
Helophorus terminassianae species	accepted (GBIF)	No associated record in PBDB
Helophorus thauma species	accepted (GBIF)	No associated record in PBDB
Helophorus theryi species	accepted (GBIF)	No associated record in PBDB
Helophorus timidus species	accepted (GBIF)	No associated record in PBDB
Helophorus trisulcatus species	accepted (GBIF)	No associated record in PBDB
Helophorus tuberculatus species	accepted (GBIF) listed (PBDB)		4	11.6 Ma Miocene	0 Ma Extant
Helophorus turca species	accepted (GBIF)	No associated record in PBDB
Helophorus uvarovi species	accepted (GBIF)	No associated record in PBDB
Helophorus villosus species	accepted (GBIF)	No associated record in PBDB
Helophorus wrootae species	accepted (GBIF)	No associated record in PBDB
Helophorus yammounensis species	accepted (GBIF)	No associated record in PBDB
Helophorus yangae species	accepted (GBIF)	No associated record in PBDB
Helophorus yixianus species	accepted (GBIF)	No associated record in PBDB
Helophorus zagrosicus species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Elophorus cinctus	GBIF	species	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : Helophorus : *Elophorus cinctus*
Kyphohelophorus Kuwert, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Kyphohelophorus*
Megahelophorus Kuwert, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Megahelophorus*
Thaumhelophorus Angus, 1995	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Thaumhelophorus*
Empleurus Hope, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Empleurus*
Gephelophorus Sharp, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Gephelophorus*
Rhopalohelophorus Kuwert, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Rhopalohelophorus*
Trichelophorus Kuwert, 1890	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Trichelophorus*
Rhopalhelophorus Kuwert, 1890	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Rhopalhelophorus*
Atractelophorus Ganglbauer, 1904	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Atractelophorus*
Orphelophorus d'Orchymont, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Orphelophorus*
Simempleurus Sharp, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Simempleurus*
Trichohelophorus Kuwert, 1888	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Trichohelophorus*
Bradempleurus Sharp, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Bradempleurus*
Cyphelophorus Seidlitz, 1888	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Cyphelophorus*
Praehelophorus Lomnicki, 1894	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Praehelophorus*
Eutrichelophorus Sharp, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Eutrichelophorus*
Helophorus Illiger, 1801	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Helophorus*
Megempleurus Sharp, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Megempleurus*
Lihelophorus Zaitzev, 1908	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Lihelophorus*
Atractohelophorus Kuwert, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Atractohelophorus*
Rhopalelophorus Kuwert, 1890	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Rhopalelophorus*
Transithelophorus Angus, 1970	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Transithelophorus*
Atracthelophorus Kuwert, 1890	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Atracthelophorus*
Atrachthelophorus Zaitzev, 1908	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Atrachthelophorus*
Elephorus Paykull, 1798	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Helophoridae : *Elephorus*

Synonymy List

Year	Name and Author
1775	Elophorus Fabricius p. 66
1792	Elophorus Fabricius p. 204
1796	Elophorus Latreille p. 8
1798	Elophorus Illiger p. 272
1802	Elophorus Latreille p. 137
1880	Helophorus Sahlberg p. 61

Year	Name and Author
1900	Helophorus Scudder p. 38
1973	Helophorus Angus
1984	Helophorus Aalto et al. p. 74
2009	Helophorus Kiselev and Nazarov
2011	Helophorus Bouchard et al. p. 153
2014	Helophorus Ponomarenko p. 1664

References

Fabricius J. C. , Systema Entomologiae, Sistens Insectorum Classes, Ordines, Genera, Species Adiectis Synonymis, Locis, Descriptionibus, Observationibus, 1-832

Fabricius J. C. , Entomologia Systematica Emendata et Aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus, Tomus I, Pars I, 1-330

Latreille P. A. , Précis des Caractères Génériques des Insectes Disposés dans un Ordre Naturel, 1-201

Illiger J. K. W. , Verzeichniss der Käfer Preussens. Entworfen von Johann Gottlieb Kugelann Apotheker in Osterode, 1-510

Latreille P. A. , Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième, 1-467

Sahlberg J. R. Bidrag till nordvestra sibiriens insektfauna, Coleoptera, insamlade under expeditionerna till obi och Jenessej 1876 och 1877: I, Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, Dryopidae, Georyssidae, Limnichidae, Heteroceridae, Staphylinidae och Micropeplidae, Kongliga Svenska Vetenskaps-Akademiens Handlingar 17 4, 1-115

Scudder S. H. Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America, Monographs of the United States Geological Survey 40, 1-148

Angus R. B. (1973) Pleistocene Helophorus (Coleoptera, Hydrophilidae) from Borislav and Starunia in the western Ukraine, with a reinterpretation of M. Lomnicki's species, description of a new Siberian species, and comparison with British Weichselian faunas, Philosophical Transactions of the Royal Society B, Biological Sciences 265, 299-326

Aalto M. M., Coope G. R., et al (1984) Late Devensian river deposits beneath the floodplain terrace of the River Thames at Abingdon, Berkshire, England, Proceedings of the Geologists Association 95, 65-79

Kiselev S. V., Nazarov V. I. (2009) Late Cenozoic insects of northern Eurasia, Paleontological Journal 43 7, 732-850

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Ponomarenko A. G. (2014) Coleoptera, in Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia, Paleontological Journal 48, 1658-1671

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

as per family - via The Interim Register of Marine and Nonmarine Genera

International Commission on Zoological Nomenclature. Opinion 1724 Helophorus Fabricius, 1775 (Insecta, Coleoptera): conserved as correct original spelling.Bulletin of Zoological Nomenclature 50, 167-168 (1993) - via Official Lists and Indexes of Names in Zoology

Hansen, Michael, 1999: null. World Catalogue of Insects, volume 2: Hydrophiloidea (s. str.) (Coleoptera). 416. - via Integrated Taxonomic Information System (ITIS)

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

Klausnitzer, B. (1994) - via Taxa Watermanagement the Netherlands (TWN)

Nilsson, A.N. (1996) - via Taxa Watermanagement the Netherlands (TWN)

Drost, M.B.P. et al. (1992) - via Taxa Watermanagement the Netherlands (TWN)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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