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Cleridae

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Description

Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences.

From Wikipedia article at https://en.wikipedia.org/wiki/Cleridae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3795	https://www.gbif.org/species/3795
PaleoBioDB ID (PBDB)	69399	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=69399

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Cleridae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Polyphaga	Emery 1886
infraorder	Cucujiformia	Lameere 1938
superfamily	Cleroidea	Latreille 1802
family	Cleridae	Latreille 1802

Common Name

checkered beetle

Scientific Name

Cleridae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Cleridae	family	belongs to Cleroidea	stated without evidence	Crowson, 1981
Cleridae	family	belongs to Coleoptera	stated with evidence	Zhang, 1989
Cleridae	family	belongs to Cleroidea	stated without evidence	Lawrence and Newton, 1995
Cleridae	family	belongs to Cleroidea	stated with evidence	Bouchard et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

25 (37%)

First Recorded Appearance

166 - 157 Ma
Jurassic

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

P. A. Latreille. 1802. Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième

Common Name(s)

Checkered Beetle, Checkered Beetles, Mierkevers, Mierenkeverachtigen, Bonte Kevers, Brokbaggar, Färgbaggar

Wikipedia

https://en.wikipedia.org/wiki/Cleridae

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Clerinae subfamily	listed (PBDB)		12	38.0 Ma Eocene	0 Ma Extant
Epiclininae subfamily	listed (PBDB)		1	166 Ma Middle Jurassic	0 Ma Extant
Hydnocerinae subfamily	listed (PBDB)		4	38.0 Ma Eocene	0 Ma Extant
Korynetinae subfamily	listed (PBDB)		7	38.0 Ma Eocene	0 Ma Extant
Tillinae subfamily	listed (PBDB)		11	38.0 Ma Eocene	0 Ma Extant
Aberrokorynetes ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Abiliella genus	accepted (GBIF)	No associated record in PBDB
Allelidea genus	accepted (GBIF)	No associated record in PBDB
Allochotes genus	accepted (GBIF)	No associated record in PBDB
Allonyx genus	accepted (GBIF)	No associated record in PBDB
Anthicoclerus genus	accepted (GBIF)	No associated record in PBDB
Aphelocerus genus	accepted (GBIF)	No associated record in PBDB
Aplemonus genus	accepted (GBIF)	No associated record in PBDB
Apteropilo genus	accepted (GBIF)	No associated record in PBDB
Araeodontia genus	accepted (GBIF)	No associated record in PBDB
Arawakis ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Aulicus genus	accepted (GBIF)	No associated record in PBDB
Avena genus	accepted (GBIF)	No associated record in PBDB
Axina genus	accepted (GBIF)	No associated record in PBDB
Barrotillus genus	accepted (GBIF)	No associated record in PBDB
Bilbotillus ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Blaesiophthalmus genus	accepted (GBIF)	No associated record in PBDB
Bogcia genus	accepted (GBIF)	No associated record in PBDB
Boschella genus	accepted (GBIF)	No associated record in PBDB
Bostrichoclerus genus	accepted (GBIF)	No associated record in PBDB
Brachycallimerus genus	accepted (GBIF)	No associated record in PBDB
Callimerus genus	accepted (GBIF)	No associated record in PBDB
Callotillus genus	accepted (GBIF)	No associated record in PBDB
Canariclerus genus	accepted (GBIF)	No associated record in PBDB
Careydon genus	accepted (GBIF)	No associated record in PBDB
Cephaloclerus genus	accepted (GBIF)	No associated record in PBDB
Chariessa genus	accepted (GBIF)	No associated record in PBDB
Cladiscus genus	accepted (GBIF)	No associated record in PBDB
Clerus genus	accepted (GBIF) listed (PBDB)				0 Ma Extant
Clytomadius genus	accepted (GBIF)	No associated record in PBDB
Colyphus genus	accepted (GBIF)	No associated record in PBDB
Coptoclerus genus	accepted (GBIF)	No associated record in PBDB
Corinthiscus genus	accepted (GBIF)	No associated record in PBDB
Corynetinus genus	accepted (GBIF)	No associated record in PBDB
Corynommadius genus	accepted (GBIF)	No associated record in PBDB
Cregya genus	accepted (GBIF)	No associated record in PBDB
Cucujocallimerus genus	accepted (GBIF)	No associated record in PBDB
Cylidrus genus	accepted (GBIF)	No associated record in PBDB
Cymatodera genus	accepted (GBIF) listed (PBDB)		2	38.0 Ma Eocene	0 Ma Extant
Cymatoderella genus	accepted (GBIF)	No associated record in PBDB
Cyrtinoclerus genus	accepted (GBIF)	No associated record in PBDB
Dasyceroclerus genus	accepted (GBIF)	No associated record in PBDB
Denops genus	accepted (GBIF)	No associated record in PBDB
Dermestoides genus	accepted (GBIF)	No associated record in PBDB
Diplocladus genus	accepted (GBIF)	No associated record in PBDB
Diplopherusa genus	accepted (GBIF)	No associated record in PBDB
Diutius genus	accepted (GBIF)	No associated record in PBDB
Egenocladiscus genus	accepted (GBIF)	No associated record in PBDB
Eleale genus	accepted (GBIF)	No associated record in PBDB
Emmepus genus	accepted (GBIF)	No associated record in PBDB
Enoclerus genus	accepted (GBIF) listed (PBDB)				0 Ma Extant
Enoplium genus	accepted (GBIF)	No associated record in PBDB
Epiclines genus	accepted (GBIF)	No associated record in PBDB
Epiphloeus genus	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	0 Ma Extant
Erymanthus genus	accepted (GBIF)	No associated record in PBDB
Eucymatodera genus	accepted (GBIF)	No associated record in PBDB
Eunatalis genus	accepted (GBIF)	No associated record in PBDB
Eurymetopum genus	accepted (GBIF)	No associated record in PBDB
Falsoorthrius genus	accepted (GBIF)	No associated record in PBDB
Flabellotilloidea genus	accepted (GBIF)	No associated record in PBDB
Gastrocentrum genus	accepted (GBIF)	No associated record in PBDB
Gracilotillus genus	accepted (GBIF)	No associated record in PBDB
Hemitrachys genus	accepted (GBIF)	No associated record in PBDB
Ichnea genus	accepted (GBIF)	No associated record in PBDB
Iontoclerus genus	accepted (GBIF)	No associated record in PBDB
Isohydnocera genus	accepted (GBIF)	No associated record in PBDB
Isolemidia genus	accepted (GBIF)	No associated record in PBDB
Kanaliella genus	accepted (GBIF)	No associated record in PBDB
Korynetes genus	accepted (GBIF)	No associated record in PBDB
Laiomorphus genus	accepted (GBIF)	No associated record in PBDB
Lasiocallimerus genus	accepted (GBIF)	No associated record in PBDB
Lebasiella genus	accepted (GBIF)	No associated record in PBDB
Lecontella genus	accepted (GBIF)	No associated record in PBDB
Lemidia genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Loedelia genus	accepted (GBIF)	No associated record in PBDB
Lyctosoma genus	accepted (GBIF)	No associated record in PBDB
Madoniella genus	accepted (GBIF)	No associated record in PBDB
Megaphloeus genus	accepted (GBIF)	No associated record in PBDB
Melanoclerus genus	accepted (GBIF)	No associated record in PBDB
Menieroclerus genus	accepted (GBIF)	No associated record in PBDB
Microclerus genus	accepted (GBIF)	No associated record in PBDB
Mimolesterus genus	accepted (GBIF)	No associated record in PBDB
Mitrandiria ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Monophylla genus	accepted (GBIF)	No associated record in PBDB
Necrobia genus	accepted (GBIF) listed (PBDB)		4	37.2 Ma Eocene	0 Ma Extant
Necrobinus genus	accepted (GBIF)	No associated record in PBDB
Neocallotillus genus	accepted (GBIF)	No associated record in PBDB
Neohydnus genus	accepted (GBIF)	No associated record in PBDB
Neorthopleura genus	accepted (GBIF)	No associated record in PBDB
Neorthrius genus	accepted (GBIF)	No associated record in PBDB
Nonalatus genus	accepted (GBIF)	No associated record in PBDB
Notostenus genus	accepted (GBIF)	No associated record in PBDB
Ohanlonella genus	accepted (GBIF)	No associated record in PBDB
Omadius genus	accepted (GBIF)	No associated record in PBDB
Onerunka genus	accepted (GBIF)	No associated record in PBDB
Onychotillus genus	accepted (GBIF)	No associated record in PBDB
Operculiphorus genus	accepted (GBIF)	No associated record in PBDB
Opetiopalpus genus	accepted (GBIF)	No associated record in PBDB
Opilo genus	accepted (GBIF)	No associated record in PBDB
Orthocladiscus genus	accepted (GBIF)	No associated record in PBDB
Orthopleuroides genus	accepted (GBIF)	No associated record in PBDB
Orthrius genus	accepted (GBIF) listed (PBDB)		2	38.0 Ma Eocene	0 Ma Extant
Pallenis genus	accepted (GBIF)	No associated record in PBDB
Parapelonides genus	accepted (GBIF)	No associated record in PBDB
Paraspinoza genus	accepted (GBIF)	No associated record in PBDB
Paratenerus genus	accepted (GBIF)	No associated record in PBDB
Paratillus genus	accepted (GBIF)	No associated record in PBDB
Parmius genus	accepted (GBIF)	No associated record in PBDB
Patuleius genus	accepted (GBIF)	No associated record in PBDB
Paupris genus	accepted (GBIF)	No associated record in PBDB
Pelonides genus	accepted (GBIF)	No associated record in PBDB
Pelonium genus	accepted (GBIF)	No associated record in PBDB
Pennasolis genus	accepted (GBIF)	No associated record in PBDB
Perilypus genus	accepted (GBIF)	No associated record in PBDB
Phlogistus genus	accepted (GBIF)	No associated record in PBDB
Phloiocopus genus	accepted (GBIF)	No associated record in PBDB
Phyllobaenus genus	accepted (GBIF) listed (PBDB)		3	38.0 Ma Eocene	0 Ma Extant
Phymatophaea genus	accepted (GBIF)	No associated record in PBDB
Placocerus genus	accepted (GBIF)	No associated record in PBDB
Placopterus genus	accepted (GBIF)	No associated record in PBDB
Plastophagus ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Plathanocera genus	accepted (GBIF)	No associated record in PBDB
Platyclerus genus	accepted (GBIF)	No associated record in PBDB
Platynoptera genus	accepted (GBIF)	No associated record in PBDB
Platytenerus genus	accepted (GBIF)	No associated record in PBDB
Plocamocera genus	accepted (GBIF)	No associated record in PBDB
Priocera genus	accepted (GBIF)	No associated record in PBDB
Prosymnus genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Protoclerus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Pseudolesterus genus	accepted (GBIF)	No associated record in PBDB
Pseudomadius genus	accepted (GBIF)	No associated record in PBDB
Pseudopallenis genus	accepted (GBIF) listed (PBDB)		4	38.0 Ma Eocene	0 Ma Extant
Pylus genus	accepted (GBIF)	No associated record in PBDB
Pylusopsis genus	accepted (GBIF)	No associated record in PBDB
Pyrrhoclerus ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Pyticeroides genus	accepted (GBIF)	No associated record in PBDB
Rhopaloclerus genus	accepted (GBIF)	No associated record in PBDB
Scrobiger genus	accepted (GBIF)	No associated record in PBDB
Sedlacekvia genus	accepted (GBIF)	No associated record in PBDB
Shartegopsis ✝ genus	accepted (GBIF) listed (PBDB)		1	152 Ma Late Jurassic	152 Ma Late Jurassic
Smudlotillus ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Spinoza genus	accepted (GBIF)	No associated record in PBDB
Stenocallimerus genus	accepted (GBIF)	No associated record in PBDB
Stenocylidrus genus	accepted (GBIF)	No associated record in PBDB
Stigmatium genus	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0 Ma Extant
Strotocera genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Tanocleria genus	accepted (GBIF)	No associated record in PBDB
Tarsostenodes genus	accepted (GBIF)	No associated record in PBDB
Tarsostenus genus	accepted (GBIF)	No associated record in PBDB
Teloclerus genus	accepted (GBIF)	No associated record in PBDB
Tenerastes genus	accepted (GBIF)	No associated record in PBDB
Teneroides genus	accepted (GBIF)	No associated record in PBDB
Teneropsis genus	accepted (GBIF)	No associated record in PBDB
Tenerus genus	accepted (GBIF)	No associated record in PBDB
Thalerocnemis genus	accepted (GBIF)	No associated record in PBDB
Thanasimodes genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Thanasimus genus	accepted (GBIF) listed (PBDB)		3	37.2 Ma Eocene	0 Ma Extant
Thriocerodes genus	accepted (GBIF)	No associated record in PBDB
Tillicera genus	accepted (GBIF)	No associated record in PBDB
Tillodenops genus	accepted (GBIF)	No associated record in PBDB
Tilloidea genus	accepted (GBIF)	No associated record in PBDB
Tillus genus	accepted (GBIF) listed (PBDB)		2	0.01 Ma Pleistocene	0 Ma Extant
Titanoclerus ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Trichodes genus	accepted (GBIF) listed (PBDB)				0 Ma Extant
Trogodendron genus	accepted (GBIF)	No associated record in PBDB
Wangweiella ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Winklerius genus	accepted (GBIF)	No associated record in PBDB
Wittmeridecus genus	accepted (GBIF)	No associated record in PBDB
Wolcottia genus	accepted (GBIF)	No associated record in PBDB
Xenorthrius genus	accepted (GBIF)	No associated record in PBDB
Xylobius genus	accepted (GBIF)	No associated record in PBDB
Zahradnikius ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Cylindus angustatus species	accepted (GBIF)	No associated record in PBDB
Elasmocylindrus tricolor species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Korynetidae	GBIF	family	Animalia : Arthropoda : Insecta : Coleoptera : Korynetidae
Trichodes stebingeri Cockerell, 1936	GBIF	species	Animalia : Arthropoda : Insecta : Coleoptera : Cleridae : *Trichodes stebingeri*
Clerus adonis Heer, 1847	GBIF	species	Animalia : Arthropoda : Insecta : Coleoptera : Cleridae : *Clerus adonis*

Synonymy List

Year	Name and Author
1802	Clerii Latreille p. 110
1847	Clerus adonis Heer p. 152 figs. Pl 5, fig 3
1865	Clerus adonis Heer p. 377
1885	Clerus adonis Scudder p. 796 fig. 1030
1936	Trichodes stebingeri Cockerell p. 5 figs. Fig 4
1981	Cleridae Crowson p. 697
1989	Cleridae Zhang p. 421
1995	Cleridae Lawrence and Newton, Jr.

Year	Name and Author
2002	Tanocleria Hong p. 130
2002	Tanocleria tenuis Hong p. 130 figs. Pl 10, fig 3; text-fig 2.6.82-87
2011	Cleridae Bouchard et al. p. 346
2015	Cleridae Robertson et al. p. 769
2016	Tanocleria Kolibá? and Huang
2016	Tanocleria tenuis Kolibá? and Huang
2016	Cleridae Webster
2019	Cleridae Gimmel et al.

References

Latreille P. A. , Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième, 1-467

Heer O. , Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Erster Theil: Käfer, 1-229

Heer O. , Die Urwelt der Schweiz, 1-622

Scudder S. H. 4. Classe Insecta. Insecten, Handbuch der Palaeontologie, I Abtheilung. Palaeozoologie 2, 747-831

Cockerell T. D. A. (1936) The Fauna of the Sunchal (or Margas Verdes) formation, Northern Argentina, American Museum Novitates 886, 1-9

Crowson R. A. (1981) The Biology of the Coleoptera, Academic Press, New York

Zhang J. F. (1989) , Fossil Insects From Shanwang, Shandong, China, 1-459

Lawrence J. F., Newton, Jr. A. F. (1995) Families and subfamilies of Coleoptera, Biology, Phylogeny and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A Crowson, 779-1006

Hong Y. C. (2002) , Amber Insects of China, 1-653

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Robertson J. A., Ślipiński A., et al (2015) Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia), Systematic Entomology 40, 745-778

Kolibáč J., Huang D. Y. (2016) The oldest known clerid fossils from the Middle Jurassic of China, with a review of Cleridae systematics (Coleoptera), Systematic Entomology 41, 808-823

Webster R. P. (2016) Checklist of the Coleoptera of New Brunswick, Canada, ZooKeys 573, 387-512

Gimmel M. L., Bocakova M., et al (2019) Comprehensive phylogeny of the Cleroidea (Coleoptera: Cucujiformia), Systematic Entomology 44, 527-558

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera

Lawrence, J. F., and A. F. Newton, Jr. / Pakaluk, James, and Stanislaw Adam Slipinski, eds., 1995: Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson, vol. 2. 779-1006. - via Integrated Taxonomic Information System (ITIS)

Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Slipinski, A.; Smith, A. (2011). Family-group names in Coleoptera (Insecta). ZooKeys. 88: 1-972. - via The Interim Register of Marine and Nonmarine Genera

Bouchard, P., Y. Bousquet, A. Davies, M. Alonso-Zarazaga, J. Lawrence, C. Lyal, A. Newton, et al., 2011: Family-group names in Coleoptera (Insecta). ZooKeys, vol. 88. 1-972. - via Integrated Taxonomic Information System (ITIS)

Opitz, Weston / Arnett, Ross H., Jr., Michael C. Thomas, Paul E. Skelley, and J. Howard Frank, eds., 2002: Family 73. Cleridae Latreille 1804. American Beetles, vol. 2: Polyphaga: Scarabaeoidea through Curculionoidea. 267-280. - via Integrated Taxonomic Information System (ITIS)

Corporaal, J. B. / Hincks, W. D., ed., 1950: null. Coleopterorum Catalogus Supplementa, Pars 23: (Editio Secunda) Cleridae. 373. - via Integrated Taxonomic Information System (ITIS)

Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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