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Agonum

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Description

Agonum is a large genus of ground beetles in the subfamily Harpalinae, tribe Platynini. They are mid-sized to smallish beetles, typically with dark metallic hues – often reddish or bronze, but sometimes black, green etc.

From Wikipedia article at https://en.wikipedia.org/wiki/Agonum, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	4988386	https://www.gbif.org/species/4988386
PaleoBioDB ID (PBDB)	193916	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=193916

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Agonum

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
subfamily	Platyninae
tribe	Platynini	Bonelli 1810
genus	Agonum	Bonelli 1810

Scientific Name

Agonum Bonelli, 1810

Name Published In

Bonelli, Franco A. 1810. Observations entomologiques. Première partie (cicindélètes et portion des carabiques). Félix Galletti, Turin.: 1-58; Tabula synoptica.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Agonum	genus	belongs to Carabiques	stated with evidence	Dejean, 1821
Agonum	genus	belongs to Carabidae	stated with evidence	Piton, 1940
Agonum	genus	belongs to Platynini	stated with evidence	Bouchard et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

18 (95%)

First Recorded Appearance

59.2 - 56.0 Ma
Paleocene

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques)

Wikipedia

https://en.wikipedia.org/wiki/Agonum

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Agonum (Europhilus) subgenus	listed (PBDB)		11	11.6 Ma Miocene	0 Ma Extant
Agonum (Olisares) subgenus	listed (PBDB)		9	0.13 Ma Pleistocene	0 Ma Extant
Agonum (Platynomicrus) subgenus	listed (PBDB)		4	0.13 Ma Pleistocene	0 Ma Extant
Agonum abnormale species	accepted (GBIF)	No associated record in PBDB
Agonum aequatum species	accepted (GBIF)	No associated record in PBDB
Agonum aeruginosum species	accepted (GBIF)	No associated record in PBDB
Agonum affine species	accepted (GBIF)	No associated record in PBDB
Agonum albicrus species	accepted (GBIF)	No associated record in PBDB
Agonum amplior species	accepted (GBIF)	No associated record in PBDB
Agonum anchomenoides species	accepted (GBIF)	No associated record in PBDB
Agonum angulifer species	accepted (GBIF)	No associated record in PBDB
Agonum antennarium species	accepted (GBIF)	No associated record in PBDB
Agonum anthracinum species	accepted (GBIF)	No associated record in PBDB
Agonum antiquum ✝ species	accepted (GBIF) listed (PBDB)		1	59.2 Ma Paleocene	59.2 Ma Paleocene
Agonum apex species	accepted (GBIF)	No associated record in PBDB
Agonum ardoisi species	accepted (GBIF)	No associated record in PBDB
Agonum arisanum species	accepted (GBIF)	No associated record in PBDB
Agonum atlantis species	accepted (GBIF)	No associated record in PBDB
Agonum atricomes species	accepted (GBIF)	No associated record in PBDB
Agonum aurelius species	accepted (GBIF)	No associated record in PBDB
Agonum azumai species	accepted (GBIF)	No associated record in PBDB
Agonum balesi species	accepted (GBIF)	No associated record in PBDB
Agonum basale species	accepted (GBIF)	No associated record in PBDB
Agonum belleri species	accepted (GBIF)	No associated record in PBDB
Agonum bellicum species	accepted (GBIF)	No associated record in PBDB
Agonum bicolor species	accepted (GBIF)	No associated record in PBDB
Agonum bilimeki species	accepted (GBIF)	No associated record in PBDB
Agonum brevicolle species	accepted (GBIF)	No associated record in PBDB
Agonum bulgani species	accepted (GBIF)	No associated record in PBDB
Agonum carbonarium species	accepted (GBIF)	No associated record in PBDB
Agonum causticum species	accepted (GBIF)	No associated record in PBDB
Agonum chalcomum species	accepted (GBIF)	No associated record in PBDB
Agonum chalconotum species	accepted (GBIF)	No associated record in PBDB
Agonum charillus species	accepted (GBIF)	No associated record in PBDB
Agonum cheni species	accepted (GBIF)	No associated record in PBDB
Agonum chinense species	accepted (GBIF)	No associated record in PBDB
Agonum chinese species	accepted (GBIF)	No associated record in PBDB
Agonum collare species	accepted (GBIF)	No associated record in PBDB
Agonum comatum species	accepted (GBIF)	No associated record in PBDB
Agonum consimile species	accepted (GBIF)	No associated record in PBDB
Agonum consimilis species	accepted (GBIF)	No associated record in PBDB
Agonum corvus species	accepted (GBIF)	No associated record in PBDB
Agonum crenistriatum species	accepted (GBIF)	No associated record in PBDB
Agonum crenulatum species	accepted (GBIF)	No associated record in PBDB
Agonum cupreum species	accepted (GBIF)	No associated record in PBDB
Agonum cupripenne species	accepted (GBIF)	No associated record in PBDB
Agonum curtipenne species	accepted (GBIF)	No associated record in PBDB
Agonum cyanopis species	accepted (GBIF)	No associated record in PBDB
Agonum cyclifer species	accepted (GBIF)	No associated record in PBDB
Agonum daimio species	accepted (GBIF)	No associated record in PBDB
Agonum darlingtoni species	accepted (GBIF)	No associated record in PBDB
Agonum deceptivum species	accepted (GBIF)	No associated record in PBDB
Agonum decorum species	accepted (GBIF)	No associated record in PBDB
Agonum deplanatum species	accepted (GBIF)	No associated record in PBDB
Agonum deuvei species	accepted (GBIF)	No associated record in PBDB
Agonum deuveiellum species	accepted (GBIF)	No associated record in PBDB
Agonum dolens species	accepted (GBIF)	No associated record in PBDB
Agonum dorsostriatum species	accepted (GBIF)	No associated record in PBDB
Agonum dreuxi species	accepted (GBIF)	No associated record in PBDB
Agonum duftschmidi species	accepted (GBIF)	No associated record in PBDB
Agonum elongatulum species	accepted (GBIF)	No associated record in PBDB
Agonum emarginatum species	accepted (GBIF)	No associated record in PBDB
Agonum ericeti species	accepted (GBIF)	No associated record in PBDB
Agonum errans species	accepted (GBIF)	No associated record in PBDB
Agonum euroum species	accepted (GBIF)	No associated record in PBDB
Agonum eurydamas species	accepted (GBIF)	No associated record in PBDB
Agonum exaratum species	accepted (GBIF)	No associated record in PBDB
Agonum excavatum species	accepted (GBIF)	No associated record in PBDB
Agonum extensicolle species	accepted (GBIF)	No associated record in PBDB
Agonum extimum species	accepted (GBIF)	No associated record in PBDB
Agonum fallax species	accepted (GBIF)	No associated record in PBDB
Agonum ferreum species	accepted (GBIF)	No associated record in PBDB
Agonum ferruginosum species	accepted (GBIF)	No associated record in PBDB
Agonum fidele species	accepted (GBIF)	No associated record in PBDB
Agonum fossiger species	accepted (GBIF)	No associated record in PBDB
Agonum fuliginosum species	accepted (GBIF)	No associated record in PBDB
Agonum galvestonicum species	accepted (GBIF)	No associated record in PBDB
Agonum gerdmuelleri species	accepted (GBIF)	No associated record in PBDB
Agonum gisellae species	accepted (GBIF)	No associated record in PBDB
Agonum gracile species	accepted (GBIF)	No associated record in PBDB
Agonum gracilipes species	accepted (GBIF)	No associated record in PBDB
Agonum graciloides species	accepted (GBIF)	No associated record in PBDB
Agonum gratiosum species	accepted (GBIF)	No associated record in PBDB
Agonum guangdongense species	accepted (GBIF)	No associated record in PBDB
Agonum harrisii species	accepted (GBIF)	No associated record in PBDB
Agonum hiranoi species	accepted (GBIF)	No associated record in PBDB
Agonum hirashimai species	accepted (GBIF)	No associated record in PBDB
Agonum holzschuhi species	accepted (GBIF)	No associated record in PBDB
Agonum humerosum species	accepted (GBIF)	No associated record in PBDB
Agonum hypocrita species	accepted (GBIF)	No associated record in PBDB
Agonum illocatum species	accepted (GBIF)	No associated record in PBDB
Agonum imitans species	accepted (GBIF)	No associated record in PBDB
Agonum impressum species	accepted (GBIF)	No associated record in PBDB
Agonum inaequale species	accepted (GBIF)	No associated record in PBDB
Agonum indiae species	accepted (GBIF)	No associated record in PBDB
Agonum iriomotense species	accepted (GBIF)	No associated record in PBDB
Agonum ishidai species	accepted (GBIF)	No associated record in PBDB
Agonum jankowskii species	accepted (GBIF)	No associated record in PBDB
Agonum japonicum species	accepted (GBIF)	No associated record in PBDB
Agonum jurecekianum species	accepted (GBIF)	No associated record in PBDB
Agonum kesharishresthae species	accepted (GBIF)	No associated record in PBDB
Agonum kucerai species	accepted (GBIF)	No associated record in PBDB
Agonum kurosonense species	accepted (GBIF)	No associated record in PBDB
Agonum kyushuense species	accepted (GBIF)	No associated record in PBDB
Agonum ladakense species	accepted (GBIF)	No associated record in PBDB
Agonum lampros species	accepted (GBIF)	No associated record in PBDB
Agonum lanei species	accepted (GBIF)	No associated record in PBDB
Agonum lanellum species	accepted (GBIF)	No associated record in PBDB
Agonum leucomerum species	accepted (GBIF)	No associated record in PBDB
Agonum liebherri species	accepted (GBIF)	No associated record in PBDB
Agonum limbaticolle species	accepted (GBIF)	No associated record in PBDB
Agonum limbatum species	accepted (GBIF)	No associated record in PBDB
Agonum longicorne species	accepted (GBIF)	No associated record in PBDB
Agonum luczoti species	accepted (GBIF)	No associated record in PBDB
Agonum lugens species	accepted (GBIF)	No associated record in PBDB
Agonum luteum species	accepted (GBIF)	No associated record in PBDB
Agonum lutulentum species	accepted (GBIF)	No associated record in PBDB
Agonum madagascariense species	accepted (GBIF)	No associated record in PBDB
Agonum mandli species	accepted (GBIF)	No associated record in PBDB
Agonum marginatum species	accepted (GBIF)	No associated record in PBDB
Agonum maruokai species	accepted (GBIF)	No associated record in PBDB
Agonum melanarium species	accepted (GBIF)	No associated record in PBDB
Agonum menetriesii species	accepted (GBIF)	No associated record in PBDB
Agonum mesostictum species	accepted (GBIF)	No associated record in PBDB
Agonum metallescens species	accepted (GBIF)	No associated record in PBDB
Agonum micans species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Agonum modestius species	accepted (GBIF)	No associated record in PBDB
Agonum moerens species	accepted (GBIF)	No associated record in PBDB
Agonum mokoto species	accepted (GBIF)	No associated record in PBDB
Agonum monachum species	accepted (GBIF)	No associated record in PBDB
Agonum mongolicum species	accepted (GBIF)	No associated record in PBDB
Agonum muelleri species	accepted (GBIF) listed (PBDB)		1	0.78 Ma Pleistocene	0 Ma Extant
Agonum muiri species	accepted (GBIF)	No associated record in PBDB
Agonum munsteri species	accepted (GBIF)	No associated record in PBDB
Agonum mutatum species	accepted (GBIF)	No associated record in PBDB
Agonum nanum species	accepted (GBIF)	No associated record in PBDB
Agonum nigriceps species	accepted (GBIF)	No associated record in PBDB
Agonum nigrum species	accepted (GBIF)	No associated record in PBDB
Agonum nipponicum species	accepted (GBIF)	No associated record in PBDB
Agonum numidicum species	accepted (GBIF)	No associated record in PBDB
Agonum nutans species	accepted (GBIF)	No associated record in PBDB
Agonum obesum species	accepted (GBIF)	No associated record in PBDB
Agonum octopunctatum species	accepted (GBIF)	No associated record in PBDB
Agonum oryctus species	accepted (GBIF)	No associated record in PBDB
Agonum pacificum species	accepted (GBIF)	No associated record in PBDB
Agonum pallipes species	accepted (GBIF)	No associated record in PBDB
Agonum palustre species	accepted (GBIF)	No associated record in PBDB
Agonum pangxiongfeii species	accepted (GBIF)	No associated record in PBDB
Agonum parextimum species	accepted (GBIF)	No associated record in PBDB
Agonum patinale species	accepted (GBIF)	No associated record in PBDB
Agonum permoestum species	accepted (GBIF)	No associated record in PBDB
Agonum persecretum species	accepted (GBIF)	No associated record in PBDB
Agonum piceolum species	accepted (GBIF)	No associated record in PBDB
Agonum piceum species	accepted (GBIF)	No associated record in PBDB
Agonum picicornoides species	accepted (GBIF)	No associated record in PBDB
Agonum placidum species	accepted (GBIF)	No associated record in PBDB
Agonum platyderum species	accepted (GBIF)	No associated record in PBDB
Agonum praetor species	accepted (GBIF)	No associated record in PBDB
Agonum propinquum species	accepted (GBIF)	No associated record in PBDB
Agonum punctiforme species	accepted (GBIF)	No associated record in PBDB
Agonum quadrimaculatum species	accepted (GBIF)	No associated record in PBDB
Agonum quadripustulatum species	accepted (GBIF)	No associated record in PBDB
Agonum quinquepuncatum species	accepted (GBIF) listed (PBDB)		1	11.6 Ma Miocene	0 Ma Extant
Agonum retractum species	accepted (GBIF)	No associated record in PBDB
Agonum rigidulum species	accepted (GBIF)	No associated record in PBDB
Agonum rotundicolle species	accepted (GBIF)	No associated record in PBDB
Agonum rougerieanum species	accepted (GBIF)	No associated record in PBDB
Agonum rufipes species	accepted (GBIF)	No associated record in PBDB
Agonum rufotestaceum species	accepted (GBIF)	No associated record in PBDB
Agonum rugaticolle species	accepted (GBIF)	No associated record in PBDB
Agonum rugicolle species	accepted (GBIF)	No associated record in PBDB
Agonum sahlbergi species	listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Agonum sataense species	accepted (GBIF)	No associated record in PBDB
Agonum scintillans species	accepted (GBIF)	No associated record in PBDB
Agonum scitulum species	accepted (GBIF)	No associated record in PBDB
Agonum scotti species	accepted (GBIF)	No associated record in PBDB
Agonum sculptipes species	accepted (GBIF)	No associated record in PBDB
Agonum scutifer species	accepted (GBIF)	No associated record in PBDB
Agonum scutiferum species	accepted (GBIF)	No associated record in PBDB
Agonum semicupreum species	accepted (GBIF)	No associated record in PBDB
Agonum sexpunctatum species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Agonum shibatai species	accepted (GBIF)	No associated record in PBDB
Agonum shibataianum species	accepted (GBIF)	No associated record in PBDB
Agonum shirahatai species	accepted (GBIF)	No associated record in PBDB
Agonum simile species	accepted (GBIF)	No associated record in PBDB
Agonum sordens species	accepted (GBIF)	No associated record in PBDB
Agonum sordidum species	accepted (GBIF)	No associated record in PBDB
Agonum speculator species	accepted (GBIF)	No associated record in PBDB
Agonum stenuiti species	accepted (GBIF)	No associated record in PBDB
Agonum straeleni species	accepted (GBIF)	No associated record in PBDB
Agonum striatopunctatum species	accepted (GBIF)	No associated record in PBDB
Agonum suavissimum species	accepted (GBIF)	No associated record in PBDB
Agonum subtruncatum species	accepted (GBIF)	No associated record in PBDB
Agonum sulcipenne species	accepted (GBIF)	No associated record in PBDB
Agonum superioris species	accepted (GBIF)	No associated record in PBDB
Agonum suturale species	accepted (GBIF)	No associated record in PBDB
Agonum sylphides species	accepted (GBIF)	No associated record in PBDB
Agonum sylphis species	accepted (GBIF)	No associated record in PBDB
Agonum tenue species	accepted (GBIF)	No associated record in PBDB
Agonum teruyai species	accepted (GBIF)	No associated record in PBDB
Agonum texanum species	accepted (GBIF)	No associated record in PBDB
Agonum thoreyi species	accepted (GBIF)	No associated record in PBDB
Agonum tokarae species	accepted (GBIF)	No associated record in PBDB
Agonum trigeminum species	accepted (GBIF)	No associated record in PBDB
Agonum triimpressum species	accepted (GBIF)	No associated record in PBDB
Agonum triseriatum species	accepted (GBIF)	No associated record in PBDB
Agonum tsushimanum species	accepted (GBIF)	No associated record in PBDB
Agonum uenoi species	accepted (GBIF)	No associated record in PBDB
Agonum variolatum species	accepted (GBIF)	No associated record in PBDB
Agonum versutum species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Agonum viduum species	accepted (GBIF)	No associated record in PBDB
Agonum viridicans species	accepted (GBIF)	No associated record in PBDB
Agonum viridicupreum species	accepted (GBIF)	No associated record in PBDB
Agonum yamatonis species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Agonocyrtes Csiki, 1931	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonocyrtes*
Agonothorax Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonothorax*
Celaenagonum Habu, 1978	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Celaenagonum*
Agonum Stephens, 1827	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonum*
Amolyntus Gistl, 1848	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amolyntus*
Agonithorax Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonithorax*
Agonocyrthes Motschoulsky, 1864	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonocyrthes*
Melanagonum Casey, 1920	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Melanagonum*
Achaetoprothorax Habu, 1978	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Achaetoprothorax*
Punctagonum Gray, 1937	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Punctagonum*
Paragonum Casey, 1920	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paragonum*
Taphranchus Casey, 1920	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Taphranchus*
Dolichodes Motschoulsky, 1864	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Dolichodes*
Agonum americanum LeConte, 1848	GBIF	species	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Agonum : *Agonum americanum*
Agonum formosum Sturm, 1826	GBIF	species	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Agonum : *Agonum formosum*
Atranodes Jedlicka, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Atranodes*
Agonium Solier, 1849	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonium*

Synonymy List

Year	Name and Author
1810	Agonum Bonelli
1821	Agonum Dejean p. 10
1940	Agonum Piton p. 171
2011	Agonum Bouchard et al. p. 140
2011	Agonum Tamutis et al.

References

Bonelli F. A. , Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques), 1-58

Dejean P. F. M. , Catalogue de la collection de Coléoptères de M. le Baron Dejean, 1-136

Piton L. (1940) , Paléontologie du Gisement Éocéne de Menat (Puy-de-Dôme) (Flore et Faune), 1-303

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Tamutis V., Tamuté B., et al (2011) A catalogue of Lithuanian beetles (Insecta, Coleoptera), ZooKeys 121, 1-494

Liebherr JK (1994) Identification of New World Agonum, review of the Mexican fauna, and description of Incagonum, new genus, from South America (Coleoptera: Carabidae: Platynini). Journal of the New York Entomological Society 102: 1-55. - via Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition.

Bousquet, Yves, and André Larochelle, 1993: Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, no. 167. 397. - via Integrated Taxonomic Information System (ITIS)

Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722. - via Integrated Taxonomic Information System (ITIS)

International Commission on Zoological Nomenclature. Opinion 1855 Agonus Bloch & Schneider, 1801 (Osteichthyes, Scorpaeniformes): conserved; and Agonidae Kirby, 1837 (Insecta, Coleoptera): spelling emended to Agonumidae, so removing the homonymy with Agonidae Swainson, 1839 (Osteichthyes, Scorpaeniformes).Bulletin of Zoological Nomenclature 53, 223-225 (1996) - via Official Lists and Indexes of Names in Zoology

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

as per family (nonmarine except where stated) - via The Interim Register of Marine and Nonmarine Genera

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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