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Enicocephalidae

Description

Enicocephalidae, also called unique-headed bugs and gnat bugs, are a family of 130 species of the suborder heteroptera. They are typically 4 mm long, and found throughout the world. They have an elongated head, constricted in places, hence their head is 'unique'.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)7880https://www.gbif.org/species/7880
PaleoBioDB ID (PBDB)213538https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=213538
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Hemiptera : Enicocephalidae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-ClareocercariaProkop et al. 2017
-Acercaria
superorderPancondylognathaYoshizawa and Leinhard 2016
-CondylognathaBörner 1904
orderHemipteraLinnaeus 1758
suborderHeteropteraLatreille 1810
-Enicocephalomorpha
familyEnicocephalidaeStal 1860
Common Nameunique-headed bug
Scientific NameEnicocephalidae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Enicocephalidaefamilybelongs to Hemipterastated with evidenceCockerell, 1917
Enicocephalidaefamilybelongs to Enicocephalomorphastated with evidenceŠtys, 2010
Status (PBDB)extant
Taxon Size (PBDB)22
Extant Size (PBDB)5 (23%)
First Recorded Appearance130 - 125 Ma
Early/Lower Cretaceous
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)T. D. A. Cockerell. 1917. Arthropods in Burmese amber. American Journal of Science, Series 4 44(263):360-368
Common Name(s) Gnat Bugs, Unique-headed Bug
Wikipediahttps://en.wikipedia.org/wiki/Enicocephalidae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Alienatinae
subfamily
listed (PBDB)420.4 Ma
Miocene
0 Ma
Extant
Disphaerocephalinae
subfamily
listed (PBDB)399.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Enicocephalinae
subfamily
listed (PBDB)156.0 Ma
Paleocene
0 Ma
Extant
Alienates
genus
accepted (GBIF)
listed (PBDB)
320.4 Ma
Miocene
0 Ma
Extant
Boreostolus
genus
accepted (GBIF)No associated record in PBDB
Brevidorsus
genus
accepted (GBIF)No associated record in PBDB
Ciucephalus
genus
accepted (GBIF)No associated record in PBDB
Disphaerocephalus
genus
accepted (GBIF)
listed (PBDB)
399.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Enicocephalinus
genus
accepted (GBIF)
listed (PBDB)
1130 Ma
Early/Lower Cretaceous
130 Ma
Early/Lower Cretaceous
Enicocephalus
genus
accepted (GBIF)
listed (PBDB)
520.4 Ma
Miocene
0 Ma
Extant
Gourlayocoris
genus
accepted (GBIF)No associated record in PBDB
Henicocephalus
genus
accepted (GBIF)No associated record in PBDB
Henschiella
genus
accepted (GBIF)No associated record in PBDB
Hoplitocoris
genus
accepted (GBIF)No associated record in PBDB
Hymenocoris
genus
accepted (GBIF)No associated record in PBDB
Lomagostus
genus
accepted (GBIF)No associated record in PBDB
Monteithostolus
genus
accepted (GBIF)No associated record in PBDB
Nesenicocephalus
genus
accepted (GBIF)No associated record in PBDB
Oncylocotis
genus
accepted (GBIF)No associated record in PBDB
Paenicotechys
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Paralienates
genus
accepted (GBIF)
listed (PBDB)
120.4 Ma
Miocene
20.4 Ma
Miocene
Phaenicocleus
genus
accepted (GBIF)No associated record in PBDB
Phthirocoris
genus
accepted (GBIF)No associated record in PBDB
Phthirostenus
genus
accepted (GBIF)No associated record in PBDB
Proboscidopirates
genus
accepted (GBIF)No associated record in PBDB
Pyrenicocephalus
genus
accepted (GBIF)
listed (PBDB)
156.0 Ma
Paleocene
56.0 Ma
Paleocene
Stenopirates
genus
accepted (GBIF)No associated record in PBDB
Systelloderes
genus
accepted (GBIF)No associated record in PBDB
Tornocrusus
genus
accepted (GBIF)No associated record in PBDB
Xenicocephalus
genus
accepted (GBIF)No associated record in PBDB
Schenchiella pauliani
species
accepted (GBIF)No associated record in PBDB

Obsolete Names

NameSourceTaxon RankTaxonomy
Enicocephalus fulvescens Westwood, 1837GBIFspeciesAnimalia : Arthropoda : Insecta : Hemiptera : Enicocephalidae : Enicocephalus fulvescens
Henicocephalidae GBIFfamilyAnimalia : Arthropoda : Insecta : Hemiptera : Henicocephalidae

Synonymy List

YearName and Author
1837Enicocephalus nasalis Hope p. 56
1837Enicocephalus fulvescens Westwood p. 23
1860Enicocephalidae Stal
1917Enicocephalidae Cockerell p. 361
2010Enicocephalidae Štys p. 456
2018Enicocephalidae Szwedo p. 112

References

Cockerell T. D. A. (1917) Arthropods in Burmese amber, American Journal of Science, Series 4 44 263, 360-368
Hope F. W. Observations on succinic insects. Part the second. Gums and resins, Transactions of the Entomological Society of London 2, 46-57
Westwood J. O. Descriptions of several new species of exotic Hemipterous insects, Transactions of the Entomological Society of London 2, 18-24
Štys P. (2010) A fossil head of an enicocephalomorphan (Hemiptera: Heteroptera) from England revisited: Identity of the first fossil species of Enicocephalidae from Europe, European Journal of Entomology 107, 455-460
Szwedo J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time, Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107, 109-128
Henry, Thomas J., and Richard C. Froeschner, eds., 1988: null. Catalog of the Heteroptera, or True Bugs, of Canada and the Continental United States. xix + 958. - via Integrated Taxonomic Information System (ITIS)
McGavin, George C., 1993: null. Bugs of the World. 192. - via Integrated Taxonomic Information System (ITIS)
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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