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Dryosauridae ✝

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Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	Not linked to a GBIF record.
PaleoBioDB ID (PBDB)	38730	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=38730

Rank

family (PBDB)

Taxonomy (PBDB)

Life : Animalia : Chordata : Ornithischia : Dryosauridae

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Triploblastica	Lankester 1877
-	Nephrozoa	Jondelius et al. 2002
-	Deuterostomia	Grobben 1908
phylum	Chordata	Haeckel 1847
subphylum	Vertebrata	Lamarck 1801
superclass	Gnathostomata	Gegenbauer 1874
-	Osteichthyes
-	Sarcopterygii	Romer 1955
subclass	Dipnotetrapodomorpha
subclass	Tetrapodomorpha
-	Tetrapoda
-	Reptiliomorpha	Säve-Söderbergh 1934
-	Anthracosauria	Säve-Söderbergh 1934
-	Batrachosauria
-	Cotylosauria
-	Amniota	Haeckel 1866
-	Sauropsida	Huxley 1864
class	Reptilia	Laurenti 1768
subclass	Eureptilia
-	Romeriida	Gauthier et al. 1988
-	Diapsida
-	Eosuchia
-	Neodiapsida	Benton 1985
-	Sauria	Gauthier 1984
-	Archosauromorpha
-	Crocopoda	Ezcurra 2016
-	Archosauriformes	Gauthier 1986
-	Eucrocopoda	Ezcurra 2016
-	Archosauria
informal	Avemetatarsalia	Benton 1999
-	Ornithodira	Gauthier 1986
-	Dinosauromorpha	Benton 1985
-	Dinosauriformes	Novas 1992
-	Dinosauria
-	Ornithischia
-	Neornithischia
-	Ornithopoda
-	Iguanodontia
-	Dryomorpha	Sereno 1986
family	Dryosauridae	Stefano 1903

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Dryosauridae	family	belongs to Proganosauria	stated without evidence	Stefano, 1903
Dryosauridae	family	belongs to Dryosauroidea	stated without evidence	Norman, 1984
Dryosauridae	family	belongs to Ornithopoda	stated without evidence	Milner and Norman, 1984
Dryosauridae	family	belongs to Ornithopoda	implied	Carroll, 1988
Dryosauridae	family	belongs to Iguanodontia	stated with evidence	Benton, 1990
Dryosauridae	family	belongs to Ornithopoda	stated with evidence	Sues and Norman, 1990
Dryosauridae	family	belongs to Ornithopoda	stated without evidence	Molnar, 1992
Dryosauridae	family	belongs to Dryomorpha	stated without evidence	Coria and Salgado, 1996
Dryosauridae	family	belongs to Iguanodontia	stated without evidence	Sereno, 1997
Dryosauridae	family	belongs to Iguanodontia	stated with evidence	Sereno, 1998
Dryosauridae	family	belongs to Dryomorpha	stated without evidence	Coria and Calvo, 2002
Dryosauridae	family	belongs to Dryomorpha	stated with evidence	Norman, 2004
Dryosauridae	family	belongs to Mesoeucrocodylia	stated without evidence	Jouve, 2005
Dryosauridae	family	belongs to Ornithopoda	stated without evidence	Chure et al., 2006
Dryosauridae	family	belongs to Iguanodontia	stated without evidence	Galton, 2007
Dryosauridae	family	belongs to Iguanodontia	stated with evidence	Butler et al., 2008
Dryosauridae	family	belongs to Dryomorpha	stated with evidence	McDonald et al., 2010
Dryosauridae	family	belongs to Iguanodontia	stated with evidence	Han et al., 2012
Dryosauridae	family	belongs to Iguanodontia	stated without evidence	Escaso et al., 2014
Dryosauridae	family	belongs to Dryomorpha	stated with evidence	Madzia et al., 2018

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

165 - 161 Ma
Jurassic

Last Recorded Appearance

70.6 - 66.0 Ma
Late/Upper Cretaceous

Environment

terrestrial (based on Ornithischia)

Motility

actively mobile (based on Ornithischia)

Diet

herbivore (based on Ornithopoda)

Reproduction

oviparous, dispersal=direct/internal,mobile (based on Ornithischia)

Ontogeny

accretion,modification of parts (based on Ornithischia)

Taphonomy

hydroxylapatite,compact or dense (based on Ornithischia)

Primary Reference (PBDB)

G. Stefano. 1903. Nuovi rettili degli strati a fosfato della Tunisia. Bollettino della Societa Geologica Italiana 22:51-80

Fossil Distribution

Subtaxa

Name	Status	Fossil Occurrences	Oldest	Youngest
Callovosaurus ✝ genus	accepted (GBIF) listed (PBDB)	1	165 Ma Middle Jurassic	165 Ma Middle Jurassic
Dinehichnus ✝ genus	listed (PBDB)	6	164 Ma Middle Jurassic	125 Ma Early/Lower Cretaceous
Dryosaurus ✝ genus	accepted (GBIF) listed (PBDB)	24	161 Ma Jurassic	151 Ma Late Jurassic
Dysalotosaurus ✝ genus	accepted (GBIF) listed (PBDB)	1	156 Ma Jurassic	156 Ma Jurassic
Elrhazosaurus ✝ genus	accepted (GBIF) listed (PBDB)	1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Eousdryosaurus ✝ genus	accepted (GBIF) listed (PBDB)	1	156 Ma Jurassic	156 Ma Jurassic
Kangnasaurus ✝ genus	accepted (GBIF) listed (PBDB)	1	101 Ma Late/Upper Cretaceous	101 Ma Late/Upper Cretaceous
Planicoxa ✝ genus	listed (PBDB)	1	129 Ma Early/Lower Cretaceous	129 Ma Early/Lower Cretaceous
Valdosaurus ✝ genus	accepted (GBIF) listed (PBDB)	9	157 Ma Jurassic	130 Ma Early/Lower Cretaceous

Synonymy List

Year	Name and Author
1903	Dryosauridae Stefano p. 54
1984	Dryosauridae Milner and Norman p. 146
1984	Dryosauridae Norman pp. 160-161 fig. 2
1988	Dryosauridae Carroll
1990	Dryosauridae Benton p. 19
1990	Dryosauridae Sues and Norman p. 500
1992	Dryosauridae Molnar p. 266
1996	Dryosauridae Coria and Salgado p. 454 fig. 13
1997	Dryosauridae Sereno p. 440 fig. 2
1998	Dryosauridae Sereno p. 61
1999	Dryosauridae Sereno p. 2138 fig. 2
2002	Dryosauridae Coria and Calvo p. 508 fig. 9

Year	Name and Author
2004	Dryosauridae Norman p. 414
2005	Dryosauridae Jouve p. 324
2006	Dryosauridae Chure et al. p. 237
2007	Dryosauridae Galton p. 38
2007	Dryosauridae Ruiz-Omeñaca et al. p. 5
2007	Dryosauridae Sánchez-Hernández et al. p. 203
2008	Dryosauridae Butler et al. p. 21 figs. 2-4
2010	Dryosauridae McDonald et al. p. 33 fig. 39
2012	Dryosauridae Han et al. p. 1391 fig. 14
2012	Dryosauridae McDonald p. 2
2014	Dryosauridae Escaso et al. p. 1103
2018	Dryosauridae Madzia et al. p. 973 fig. 4

References

Stefano G. (1903) Nuovi rettili degli strati a fosfato della Tunisia, Bollettino della Societa Geologica Italiana 22, 51-80

Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698

Benton M. J. (1990) Origin and interrelationships of dinosaurs, The Dinosauria, D. B. Weishampel, P. Dodson, & H. Osmólska (editors), University of California Press, Berkeley, 11-30

Coria R. A., Salgado L. (1996) A basal iguanodontian (Ornithischia: Ornithopoda) from the Late Cretaceous of South America, Journal of Vertebrate Paleontology 16 3, 445-457

Sereno P. C. (1997) The origin and evolution of dinosaurs, Annual Review of Earth and Planetary Sciences 25, 435-489

Sereno P. C. (1998) A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria, Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 210 1, 41-83

Sereno P. C. (1999) The evolution of dinosaurs, Science 284, 2137-2147

Coria R. A., Calvo J. O. (2002) A new iguanodontian ornithopod from Neuquén Basin, Patagonia, Argentina, Journal of Vertebrate Paleontology 22 3, 503-509

Norman D. B. (2004) Basal Iguanodontia, The Dinosauria (2nd edition). University of California Press, Berkeley, 413-437

Jouve S. (2005) A new description of the skull of Dyrosaurus phosphaticus (Thomas, 1893) (Mesoeucrocodylia: Dyrosauridae) from the Lower Eocene of North Africa, Canadian Journal of Earth Sciences 42, 323-337

Chure D. J., Litwin R., et al (2006) The fauna and flora of the Morrison Formation: 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36, 233-249

Galton P. M. (2007) Teeth of ornithischian dinosaurs (mostly Ornithopoda) from the Morrison Formation (Upper Jurassic) of the western United States, Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, 17-47

Ruiz-Omeñaca J. I., Pereda Suberbiola X., et al (2007) Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England, Horns and Beaks: Ceratopsian and Ornithischian Dinosaurs, 3-16

Sánchez-Hernández B., Benton M. J., et al (2007) Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain, Palaeogeography, Palaeoclimatology, Palaeoecology 249, 180-215 doi:10.1016/j.palaeo.2007.01.009

Butler R. J., Upchurch P., et al (2008) The phylogeny of ornithischian dinosaurs, Journal of Systematic Palaeontology 6 1, 1-40 doi:10.1017/S1477201907002271

McDonald A. T., Kirkland J. I., et al (2010) New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs, PLoS One 5 11, e14075:1-35 doi:10.1371/journal.pone.0014075

Han F.-L., Barrett P. M., et al (2012) Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China, Journal of Vertebrate Paleontology 32 6, 1370-1395 doi:10.1080/02724634.2012.694385

McDonald A. T. (2012) Phylogeny of basal iguanodonts (Dinosauria: Ornithischia): an update, PLoS ONE 7 5, e36745:1-5 doi:10.1371/journal.pone.0036745

Escaso F., Ortega F., et al (2014) A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal, Journal of Vertebrate Paleontology 34 5, 1102-1112 doi:10.1080/02724634.2014.849715

Madzia D., Boyd C. A., et al (2018) A basal ornithopod dinosaur from the Cenomanian of the Czech Republic, Journal of Systematic Palaeontology 16 11, 967-979 doi:10.1080/14772019.2017.1371258

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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