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Pachycephalosauridae ✝

Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)Not linked to a GBIF record.
PaleoBioDB ID (PBDB)38783https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=38783
Rankfamily (PBDB)
Taxonomy (PBDB)Life : Animalia : Chordata : Ornithischia : Pachycephalosauridae
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-TriploblasticaLankester 1877
-NephrozoaJondelius et al. 2002
-DeuterostomiaGrobben 1908
phylumChordataHaeckel 1847
subphylumVertebrataLamarck 1801
superclassGnathostomataGegenbauer 1874
-Osteichthyes
-SarcopterygiiRomer 1955
subclassDipnotetrapodomorpha
subclassTetrapodomorpha
-Tetrapoda
-ReptiliomorphaSäve-Söderbergh 1934
-AnthracosauriaSäve-Söderbergh 1934
-Batrachosauria
-Cotylosauria
-AmniotaHaeckel 1866
-SauropsidaHuxley 1864
classReptiliaLaurenti 1768
subclassEureptilia
-RomeriidaGauthier et al. 1988
-Diapsida
-Eosuchia
-NeodiapsidaBenton 1985
-SauriaGauthier 1984
-Archosauromorpha
-CrocopodaEzcurra 2016
-ArchosauriformesGauthier 1986
-EucrocopodaEzcurra 2016
-Archosauria
informalAvemetatarsaliaBenton 1999
-OrnithodiraGauthier 1986
-DinosauromorphaBenton 1985
-DinosauriformesNovas 1992
-Dinosauria
-Ornithischia
-Neornithischia
-ClypeodontaNorman 2014
-CerapodaSereno 1986
-MarginocephaliaSereno 1986
-Pachycephalosauria
familyPachycephalosauridaeSternberg 1945
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceSternberg, 1945
Pachycephalosauridaefamilybelongs to OrnithopodaimpliedRomer, 1956
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceParsch, 1963
Pachycephalosauridaefamilybelongs to Ornithischiastated with evidenceTatarinov, 1964
Pachycephalosauridaefamilybelongs to Ornithopodastated with evidenceEstes, 1964
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceKuhn, 1964
Pachycephalosauridaefamilybelongs to OrnithopodaimpliedRomer, 1966
Pachycephalosauridaefamilybelongs to Ornithopodastated with evidenceKuhn, 1966
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceSwinton, 1970
Pachycephalosauridaefamilybelongs to Brachypodastated without evidenceThulborn, 1974
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceGalton, 1974
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceMaryanska and Osmolska, 1974
Pachycephalosauridaefamilybelongs to Brachypodastated without evidenceThulborn, 1977
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceRozhdestvensky, 1977
Pachycephalosauridaefamilybelongs to Ornitophodastated without evidenceBonaparte, 1978
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceDong, 1978
Pachycephalosauridaefamilybelongs to OrnithopodaimpliedBenton, 1979
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceSues and Taquet, 1979
Pachycephalosauridaefamilybelongs to Pachycephalosauroideastated without evidenceZhao, 1983
Pachycephalosauridaefamilybelongs to Ornithopodastated without evidenceNorman, 1984
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceRussell, 1984
Pachycephalosauridaefamilybelongs to Orbitoscutastated without evidenceBakker, 1986
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceSues and Galton, 1987
Pachycephalosauridaefamilybelongs to PachycephalosauriaimpliedCarroll, 1988
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceMaryanska, 1990
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceDong, 1992
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceSereno, 1998
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidenceGoodwin et al., 1998
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceSereno, 1999
Pachycephalosauridaefamilybelongs to Ornithischiastated without evidenceEberth et al., 2001
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated without evidencePeng et al., 2001
Pachycephalosauridaefamilybelongs to Homalocephaloideastated with evidenceMaryanska et al., 2004
Pachycephalosauridaefamilybelongs to Ornithischiastated with evidenceSullivan, 2006
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceButler et al., 2008
Pachycephalosauridaefamilybelongs to Ornithischiastated without evidenceJasinski and Sullivan, 2011
Pachycephalosauridaefamilybelongs to Pachycephalosauriastated with evidenceHan et al., 2012
Pachycephalosauridaefamilybelongs to Ornithischiastated without evidenceEvans et al., 2013
Status (PBDB)extinct
Taxon Size (PBDB)27
First Recorded Appearance101 - 84 Ma
Late/Upper Cretaceous
Last Recorded Appearance70.6 - 66.0 Ma
Late/Upper Cretaceous
Environmentterrestrial (based on Ornithischia)
Motilityactively mobile (based on Ornithischia)
Dietherbivore (based on Ornithischia)
Reproductionoviparous, dispersal=direct/internal,mobile (based on Ornithischia)
Ontogenyaccretion,modification of parts (based on Ornithischia)
Taphonomyhydroxylapatite,compact or dense (based on Ornithischia)
Primary Reference (PBDB)C. M. Sternberg. 1945. Pachycephalosauridae proposed for dome-headed dinosaurs, Stegoceras lambei, n. sp., described. Journal of Paleontology 19(5):534-538

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Domocephalinae
subfamily
listed (PBDB)
Pachycephalosaurinae
subfamily
listed (PBDB)
Acrotholus
genus
listed (PBDB)185.8 Ma
Late/Upper Cretaceous
85.8 Ma
Late/Upper Cretaceous
Amtocephale
genus
listed (PBDB)1101 Ma
Late/Upper Cretaceous
101 Ma
Late/Upper Cretaceous
Colepiocephale
genus
listed (PBDB)483.5 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Hanssuesia
genus
listed (PBDB)283.6 Ma
Late/Upper Cretaceous
83.6 Ma
Late/Upper Cretaceous
Prenocephale
genus
listed (PBDB)1183.6 Ma
Late/Upper Cretaceous
72.1 Ma
Late/Upper Cretaceous
Stegoceras
genus
listed (PBDB)4183.6 Ma
Late/Upper Cretaceous
72.1 Ma
Late/Upper Cretaceous
Tylocephale
genus
listed (PBDB)183.6 Ma
Late/Upper Cretaceous
83.6 Ma
Late/Upper Cretaceous
Pachycephalosaurini
unranked clade
listed (PBDB)3183.5 Ma
Late/Upper Cretaceous
70.6 Ma
Late/Upper Cretaceous

Synonymy List

YearName and Author
1945Pachycephalosauridae Sternberg pp. 535-536
1953Troodon bexelli Bohlin p. 32
1956Pachycephalosauridae Romer p. 631
1963Pachycephalosauridae Parsch p. 4
1964Pachycephalosauridae Estes p. 144
1964Pachycephalosauridae Kuhn p. 30
1964Stegoceras bexelli Kuhn p. 32
1964Pachycephalosauridae Tatarinov p. 588
1966Pachycephalosauridae Kuhn p. 103
1966Pachycephalosauridae Romer p. 370
1970Pachycephalosauridae Swinton p. 200
1971Pachycephalosauridae Galton p. 40
1974Pachycephalosauridae Galton p. 1064 fig. 6
1974Pachycephalosauridae Maryanska and Osmolska p. 50
1974Pachycephalosauridae Thulborn p. 173
1977Pachycephalosauridae Rozhdestvensky p. 114
1977Pachycephalosauridae Thulborn p. 726 fig. 1
1978Pachycephalosauridae Bonaparte p. 331
1978Pachycephalosauridae Dong p. 225
1979Pachycephalosauridae Benton p. 143
1979Pachycephalosauridae Sues and Taquet
1979Gravitholus Wall and Galton p. 1180
1979Pachycephalosauridae Wall and Galton p. 1185
1980Pachycephalosauridae Bonaparte p. 82
1980Troodon bexell Chao
1980Gravitholus Currie pp. 29-30
1980Pachycephalosauridae Ostrom p. 22
1981Pachycephalosauridae McIntosh p. 35
1983Gravitholus Dodson p. 95
1983Pachycephalosauridae Zhao p. 301
1984Pachycephalosauridae Norman pp. 160-162 fig. 2
1984Gravitholus Russell p. 25
1984Pachycephalosauridae Russell p. 29
YearName and Author
1985Pachycephalosauridae Cooper p. 291
1986Pachycephalosauridae Bakker p. 461
1986Gravitholus Clemens p. 80
1986Tholocephalidae Sereno
1986Stegoceras bexell Zhao p. 71
1987Gravitholus Sues and Galton
1987Pachycephalosauridae Sues and Galton
1988Gravitholus Carroll
1988Pachycephalosauridae Carroll
1990Gravitholus Maryanska p. 566
1990Pachycephalosauridae Maryanska p. 566
1992Pachycephalosauridae Dong p. 166
1997Troodon bexelli Mateer p. 662
1997Pachycephalosauridae Sereno p. 443
1997Pachycephalosauridae Sues p. 512
1998Pachycephalosauridae Goodwin et al. p. 363
1998Pachycephalosauridae Sereno p. 62
1999Pachycephalosauridae Sereno p. 2138 fig. 2
2000Pachycephalosauridae Sereno p. 482
2001Pachycephalosauridae Eberth et al. p. 57
2001Pachycephalosauridae Peng et al. p. 36
2003Pachycephalosauridae Williamson and Carr fig. 11
2004Gravitholus Maryanska et al. p. 465
2004Pachycephalosauridae Maryanska et al. p. 465
2006Pachycephalosauridae Sullivan p. 349
2008Pachycephalosauridae Butler et al. pp. 22-23 figs. 2-4
2010Texacephale Longrich et al.
2010Texacephale langstoni Longrich et al. p. 276 figs. 3-5
2011Pachycephalosauridae Jasinski and Sullivan p. 202
2011Pachycephalosauridae Watabe et al. p. 490
2012Pachycephalosauridae Han et al. p. 1391 fig. 14
2013Pachycephalosauridae Evans et al. p. 2
2013Texacephale langstoni Longrich et al. p. 24

References

Sternberg C. M. (1945) Pachycephalosauridae proposed for dome-headed dinosaurs, Stegoceras lambei, n. sp., described, Journal of Paleontology 19 5, 534-538
Bohlin B. (1953) VI. Vertebrate Palaeontology 6. Fossil reptiles from Mongolia and Kansu, Reports from the Scientific Expedition to the North-western Provinces of China under Leadership of Dr. Sven Hedin. The Sino-Swedish Expedition Publication 37, 1-113
Romer A. S. (1956) , Osteology of the Reptiles, University of Chicago Press, 1-772
Estes R. (1964) Fossil vertebrates from the Late Cretaceous Lance Formation, eastern Wyoming, University of California Publications in Geological Sciences 49, 1-187
Tatarinov L. P. (1964) Nadotryad Dinosauria. Dinozavry [Superorder Dinosauria. Dinosaurs], Osnovy Paleontologii [Fundamentals of Paleontology] 12, 523-589
Kuhn O. W. M. (1966) , Die Reptilien. System und Stammesgeschichte [The Reptiles. Systematics and Phylogeny.], 1-154
Romer A. S. (1966) , Vertebrate Paleontology, 3rd edition, 1-468
Swinton W. E. (1970) , The Dinosaurs, Wiley-Interscience, New York, 1-331
Galton P. M. (1971) A primitive dome-headed dinosaur (Ornithischia: Pachycephalosauridae) from the Lower Cretaceous of England and the function of the dome of pachycephalosaurids, Journal of Paleontology 45 1, 40-47
Galton P. M. (1974) Notes on Thescelosaurus, a conservative ornithopod dinosaur from the Upper Cretaceous of North America, with comments on ornithopod classification, Journal of Paleontology 48 5, 1048-1067
Maryanska T., Osmolska H. (1974) Pachycephalosauria, a new suborder of ornithischian dinosaurs, Palaeontologia Polonica 30, 45-102
Thulborn R. A. (1974) A new heterodontosaurid dinosaur (Reptilia: Ornithischia) from the Upper Triassic red beds of Lesotho, Zoological Journal of the Linnean Society 55, 151-175
Rozhdestvensky A. K. (1977) The study of dinosaurs in Asia, Journal of the Palaeontological Society of India 20, 102-119
Bonaparte J. F. (1978) El Mesozoico de America de Sur y sus Tetrapodos [The Mesozoic of South America and its tetrapods], Opera Lilloana 26, 1-596
Dong Z. (1978) [A new genus of Pachycephalosauria from Laiyang, Shantung], Vertebrata PalAsiatica 16 4, 225-228
Wall W. P., Galton P. M. (1979) Notes on pachycephalosaurid dinosaurs (Reptilia: Ornithischia) from North America, with comments on their status as ornithopods, Canadian Journal of Earth Sciences 16, 1176-1186
Currie P. J. (1980) Mesozoic vertebrate life in Alberta and British Columbia, Mesozoic Vertebrate Life 1, 27-40
Ostrom J. H. (1980) The evidence for endothermy in dinosaurs, In E. C. Olson & R. D. K. Thomas (eds.), A Cold Look at the Warm-Blooded Dinosaurs. AAAS Selected Symposium 28. Westview Press, Boulder, CO, 15-54
McIntosh J. S. (1981) Annotated catalogue of the dinosaurs (Reptilia, Archosauria) in the collections of the Carnegie Museum of Natural History, Bulletin of Carnegie Museum of Natural History 18, 1-67
Zhao X. (1983) Phylogeny and evolutionary stages of Dinosauria, Acta Palaeontologica Polonica 28 1-2, 295-306
Russell D. A. (1984) A check list of the families and genera of North American dinosaurs, Syllogeus 53, 1-35
Bakker R. T. (1986) , The Dinosaur Heresies. William Morrow and Company, Inc, New York, 1-481
Sereno P. C. (1986) Phylogeny of the bird-hipped dinosaurs (order Ornithischia), National Geographic Research 2 2, 234-256
Zhao X. (1986) [Reptiles], Ching-kuo Ti Pao o Hsi [The Cretaceous System of China. The Stratigraphy of China] 12, 67-73
Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698
Dong Z. (1992) Dinosaurian Faunas of China, China Ocean Press, Beijing, 1-188
Sereno P. C. (1997) The origin and evolution of dinosaurs, Annual Review of Earth and Planetary Sciences 25, 435-489
Goodwin M. B., Buccholtz E. A., et al (1998) Cranial anatomy and diagnosis of Stygimoloch spinifer (Ornithischia: Pachycephalosauria) with comments on cranial display structures in agonistic behavior, Journal of Vertebrate Paleontology 18 2, 363-375
Sereno P. C. (1998) A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria, Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 210 1, 41-83
Sereno P. C. (1999) The evolution of dinosaurs, Science 284, 2137-2147
Sereno P. C. (2000) The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia, The Age of Dinosaurs in Russia and Mongolia, 480-516
Eberth D. A., Currie P. J., et al (2001) Alberta's dinosaurs and other fossil vertebrates: Judith River and Edmonton groups (Campanian-Maastrichtian), In C. L. Hill (ed), Society of Vertebrate Paleontology, 61st Annual Meeting, Bozeman. Guidebook for the Field Trips: Mesozoic and Cenozoic Paleontology in the Western Plains and Rocky Mountains, Museum of the Rockies Occasional Paper 3, 49-75
Peng J., Russell A. P., et al (2001) Vertebrate microsite assemblages (exclusive of mammals) from the Foremost and Oldman Formations of the Judith River Group (Campanian) of southeastern Alberta: an illustrated guide, Provincial Museum of Alberta, Natural History Occasional Paper 25, 1-54
Williamson T. E., Carr T. D. (2003) A new genus of derived pachycephalosaurian from western North America, Journal of Vertebrate Paleontology 22 4, 779-801
Sullivan R. M. (2006) A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia), Late Cretaceous Vertebrates from the Western Interior. New Mexico Museum of Natural History and Science Bulletin 35, 348-365
Butler R. J., Upchurch P., et al (2008) The phylogeny of ornithischian dinosaurs, Journal of Systematic Palaeontology 6 1, 1-40 doi:10.1017/S1477201907002271
Longrich N. R., Sankey J., et al (2010) Texacephale langstoni, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA, Cretaceous Research 31, 274-284 doi:10.1016/j.cretres.2009.12.002
Jasinski S. E., Sullivan R. M. (2011) Re-evaluation of pachycephalosaurids from the Fruitland-Kirtland transition (Kirtlandian, late Campanian), San Juan Basin, New Mexico, with a description of a new species of Stegoceras and a reassessment of Texacephale langstoni, Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin 53, 202-215
Watabe M., Tsogtbaatar K., et al (2011) A new pachycephalosaurid from the Baynshire Formation (Cenomanian-late Santonian), Gobi Desert, Mongolia, Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin 53, 489-497
Han F.-L., Barrett P. M., et al (2012) Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China, Journal of Vertebrate Paleontology 32 6, 1370-1395 doi:10.1080/02724634.2012.694385
Evans D. C., Schott R. K., et al (2013) The oldest North American pachycephalosaurid and the hidden diversity of small-bodied ornithischian dinosaurs, Nature Communications 4, 1828 doi:10.1038/ncomms2749
Longrich N. R., Barnes K., et al (2013) Caenagnathidae from the upper Campanian Aguja Formation of west Texas, and a revision of the Caenagnathinae, Bulletin of the Peabody Museum of Natural History 54 1, 23-49 doi:http://dx.doi.org/10.3374/014.054.0102
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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