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Ithonidae

Description

Ithonidae, commonly called moth lacewings and giant lacewings, is a small family of winged insects of the insect order Neuroptera. The family contains a total of ten living genera, and twelve extinct genera described from fossils. The modern Ithonids have a notably disjunct distribution, while the extinct genera had a more global range. The family is considered one of the most primitive living neuropteran families. Ithonids may be phytophagous. The family has been expanded twice, first to include the genus Rapisma, formerly placed in the monotypic family Rapismatidae, and then in 2010 to include the genera that had been placed into the family Polystoechotidae. Both Rapismatidae and Polystoechotidae have been shown to nest into Ithonidae.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)5012https://www.gbif.org/species/5012
PaleoBioDB ID (PBDB)193501https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=193501
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Neuroptera : Ithonidae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-Holometabola
-Aparaglossata
superorderNeuropterida
-EidoneuropteraEngel et al. 2018
orderNeuropteraLinnaeus 1758
-EuneuropteraEngel et al. 2018
-NeoneuropteraEngel et al. 2018
-GeoneuropteraEngel et al. 2018
superfamilyIthonoideaNewman 1853
familyIthonidaeNewman 1853
Common Namemoth lacewing
Scientific NameIthonidae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Ithonidaefamilybelongs to Ithonoideastated with evidenceEngel and Grimaldi, 2008
Ithonidaefamilybelongs to Neuropterastated without evidenceMakarkin and Archibald, 2009
Ithonidaefamilybelongs to Ithonoideastated with evidenceEngel et al., 2018
Status (PBDB)extant
Taxon Size (PBDB)19
Extant Size (PBDB)1 (5.3%)
First Recorded Appearance166 - 157 Ma
Jurassic
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)V. N. Makarkin and S. B. Archibald. 2009. A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America. Zootaxa 2063:55-63
Common Name(s) Giant Lacewings, Moth Lacewings, Mothlike Lacewings, Ithonid Lacewings, Moth-like Lacewings, Moth Lacewing
Wikipediahttps://en.wikipedia.org/wiki/Ithonidae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Adamsiana
genus
accepted (GBIF)No associated record in PBDB
Allorapisma
genus
accepted (GBIF)
listed (PBDB)
256.0 Ma
Paleocene
56.0 Ma
Paleocene
Burmithone
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Cratovoluptia
genus
accepted (GBIF)
listed (PBDB)
1122 Ma
Early/Lower Cretaceous
122 Ma
Early/Lower Cretaceous
Elektrithone
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Fontecilla
genus
accepted (GBIF)No associated record in PBDB
Guithone
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Ithone
genus
accepted (GBIF)No associated record in PBDB
Jurapolystoechotes
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Kirgisella
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Lasiosmylus
genus
accepted (GBIF)
listed (PBDB)
4125 Ma
Early/Lower Cretaceous
125 Ma
Early/Lower Cretaceous
Megalithone
genus
accepted (GBIF)No associated record in PBDB
Meilingius
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Mesopolystoechus
genus
accepted (GBIF)
listed (PBDB)
2191 Ma
Early Jurassic
166 Ma
Middle Jurassic
Narodona
genus
accepted (GBIF)No associated record in PBDB
Oliarces
genus
accepted (GBIF)No associated record in PBDB
Osmyloides
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Palaeopsychops
genus
accepted (GBIF)
listed (PBDB)
2856.0 Ma
Paleocene
56.0 Ma
Paleocene
Paleopterocalla
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Panfilovdvia
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Platystoechotes
genus
accepted (GBIF)No associated record in PBDB
Polystoechotes
genus
accepted (GBIF)
listed (PBDB)
0 Ma
Extant
Polystoechotites
genus
accepted (GBIF)
listed (PBDB)
656.0 Ma
Paleocene
37.2 Ma
Eocene
Principiala
genus
accepted (GBIF)
listed (PBDB)
3130 Ma
Early/Lower Cretaceous
122 Ma
Early/Lower Cretaceous
Puripolystoechotes
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Rapisma
genus
accepted (GBIF)No associated record in PBDB
Ricaniella
genus
accepted (GBIF)
listed (PBDB)
156.0 Ma
Paleocene
56.0 Ma
Paleocene
Varnia
genus
accepted (GBIF)No associated record in PBDB

Obsolete Names

NameSourceTaxon RankTaxonomy
Polystoechotidae GBIFfamilyAnimalia : Arthropoda : Insecta : Neuroptera : Polystoechotidae

Synonymy List

YearName and Author
1853Ithonidae Newman
2008Ithonidae Engel and Grimaldi
2009Ithonidae Makarkin and Archibald
2018Ithonidae Engel et al.
1853Ithonidae Newman
2008Ithonidae Engel and Grimaldi
2009Ithonidae Makarkin and Archibald
2018Ithonidae Engel et al.

References

Makarkin V. N., Archibald S. B. (2009) A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America, Zootaxa 2063, 55-63
Engel M. S., Grimaldi D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta), Nova Supplementa Entomologica 20, 1-86
Engel M. S., Winterton S. L., et al (2018) Phylogeny and evolution of Neuropterida: where have wings of lace taken us?, Annual Review of Entomology 63, 531-551
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
Winterton, Shaun L., and Vladimir N. Makarkin, 2010: Phylogeny of Moth Lacewings and Giant Lacewings (Neuroptera: Ithonidae, Polystoechotidae) Using DNA Sequence Data, Morphology, and Fossils. Annals of the Entomological Society of America, vol. 103, no. 4. 511-522. - via Integrated Taxonomic Information System (ITIS)
Penny, Norman D., Philip A. Adams, and Lionel A. Stange, 1997: Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico. Proceedings of the California Academy of Sciences, vol. 50, no. 3. 39-114. - via Integrated Taxonomic Information System (ITIS)
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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