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Arthropoda

Photos

Photo Gallery

Eurypterus sp.

Buffalo, Erie County, New York, USA

Euproops sp.

Mazon Creek, Will County, Illinois, USA

Kainops invius

Coal County, Oklahoma, USA

Eurypterus sp.

Buffalo, Erie County, New York, USA

Euproops sp.

Mazon Creek, Will County, Illinois, USA

Kainops invius

Coal County, Oklahoma, USA

Eurypterus sp.

Buffalo, Erie County, New York, USA

Euproops sp.

Mazon Creek, Will County, Illinois, USA

Description

An arthropod (from Greek ἄρθρον arthron, "joint" and πούς pous, "foot" (gen. ποδός)) is an invertebrate animal having an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Euarthropoda, which includes insects, arachnids, myriapods, and crustaceans. The term Arthropoda as originally proposed refers to a proposed grouping of Euarthropods and the phylum Onychophora.

From Wikipedia article at https://en.wikipedia.org/wiki/Arthropoda, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	54	https://www.gbif.org/species/54
PaleoBioDB ID (PBDB)	18891	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=18891

Rank

phylum

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829

Common Name

arthropod

Scientific Name

Arthropoda

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Arthropoda	phylum	belongs to Metazoa	stated without evidence	Shoshani et al., 1989
Arthropoda	phylum	belongs to Panarthropoda	stated without evidence	Zrzavy et al., 1998
Arthropoda	phylum	belongs to Metazoa	second hand	Sepkoski, 2002
Arthropoda	phylum	belongs to Ecdysozoa	stated without evidence	Edgecombe et al., 2011
Arthropoda	phylum	belongs to Panarthropoda	stated without evidence	Nielsen, 2012

Status (PBDB)

extant

Taxon Size (PBDB)

71,493

Extant Size (PBDB)

13,479 (19%)

First Recorded Appearance

635 - 541 Ma
Neoproterozoic

Motility

actively mobile

Primary Reference (PBDB)

J. J. Sepkoski, Jr. 2002. A compendium of fossil marine animal genera. Bulletins of American Paleontology 363:1-560

Common Name(s)

Arthropods, Arthropodes, Artrópode, Arthropod, Leddjur, Gliederfüßer, Geleedpotigen, Arthropoden, Членистоногі, 節足動物

Wikipedia

https://en.wikipedia.org/wiki/Arthropoda

External Images

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Arachnomorpha subphylum	listed (PBDB)	Trilobite-like	119	520 Ma Cambrian	0 Ma Extant
Chelicerata subphylum	listed (PBDB)		4,074	520 Ma Cambrian	0 Ma Extant
Mandibulata subphylum	listed (PBDB)		133,726	541 Ma Cambrian	0 Ma Extant
Proschizoramia superclass	listed (PBDB)		35	520 Ma Cambrian	513 Ma Cambrian
Arachnida class	accepted (GBIF) listed (PBDB)		3,515	439 Ma Silurian	0 Ma Extant
Arachnoidea ✝ class	listed (PBDB)
Artiopoda ✝ class	listed (PBDB)	Artiopod	38	520 Ma Cambrian	501 Ma Cambrian
Branchiopoda class	accepted (GBIF) listed (PBDB)	Branchiopod	164	520 Ma Cambrian	0 Ma Extant
Cephalocarida class	accepted (GBIF) listed (PBDB)		1	501 Ma Cambrian	0 Ma Extant
Chilopoda class	accepted (GBIF) listed (PBDB)	Centipede	52	423 Ma Silurian	0 Ma Extant
Dinocarida ✝ class	listed (PBDB)		1	513 Ma Cambrian	513 Ma Cambrian
Diplopoda class	accepted (GBIF) listed (PBDB)	Millipede	338	433 Ma Silurian	0 Ma Extant
Entognatha class	accepted (GBIF)	No associated record in PBDB
Hexanauplia class	accepted (GBIF)	No associated record in PBDB
Insecta class	accepted (GBIF) listed (PBDB)	Insect	47,122	372 Ma Late/Upper Devonian	0 Ma Extant
Malacostraca class	accepted (GBIF) listed (PBDB)		6,175	530 Ma Cambrian	0 Ma Extant
Marrelomorpha class	listed (PBDB)
Maxillopoda class	accepted (GBIF) listed (PBDB)		1,054	433 Ma Silurian	0 Ma Extant
Merostomata class	accepted (GBIF)	No associated record in PBDB
Merostomoidea class	listed (PBDB)		1	478 Ma Early/Lower Ordovician	478 Ma Early/Lower Ordovician
Myriapoda class	listed (PBDB)		415	433 Ma Silurian	0 Ma Extant
Ostracoda class	accepted (GBIF) listed (PBDB)	Ostracod	26,236	541 Ma Cambrian	0 Ma Extant
Paratrilobita class	listed (PBDB)		5	635 Ma Neoproterozoic	635 Ma Neoproterozoic
Pauropoda class	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Pycnogonida class	accepted (GBIF) listed (PBDB)	Sea Spider	4	433 Ma Silurian	0 Ma Extant
Remipedia class	accepted (GBIF) listed (PBDB)		1	331 Ma Carboniferous	0 Ma Extant
Scorpionida class	listed (PBDB)	Scorpion	6	443 Ma Silurian	0 Ma Extant
Symphyla class	accepted (GBIF) listed (PBDB)		5	99.6 Ma Late/Upper Cretaceous	0 Ma Extant
Trilobita ✝ class	accepted (GBIF) listed (PBDB)	Trilobite	52,454	541 Ma Cambrian	254 Ma Permian
Euthycarcinoidea ✝ subclass	listed (PBDB)		6	411 Ma Early/Lower Devonian	247 Ma Early/Lower Triassic
Bradoriida ✝ order	listed (PBDB)		274	541 Ma Cambrian	41.3 Ma Eocene
Fuxianhuiida ✝ order	listed (PBDB)		25	541 Ma Cambrian	516 Ma Cambrian
Gigantopleurida ✝ order	listed (PBDB)
Mystacocaridida order	accepted (GBIF)	No associated record in PBDB
Nektaspida ✝ order	listed (PBDB)		139	520 Ma Cambrian	445 Ma Late/Upper Ordovician
Orstenocarida ✝ order	listed (PBDB)		1	501 Ma Cambrian	501 Ma Cambrian
Radiodonta ✝ order	listed (PBDB)		140	541 Ma Cambrian	513 Ma Cambrian
Skaracarida ✝ order	listed (PBDB)		2	501 Ma Cambrian	501 Ma Cambrian
Xandarellida ✝ order	listed (PBDB)		1	520 Ma Cambrian	520 Ma Cambrian
Aglaspididae family	accepted (GBIF)	No associated record in PBDB
Amplectobeluidae family	accepted (GBIF)	No associated record in PBDB
Anomalocarididae family	accepted (GBIF)	No associated record in PBDB
Archidesmidae family	accepted (GBIF)	No associated record in PBDB
Arrhopalitida family	accepted (GBIF)	No associated record in PBDB
Arthropleuridae family	accepted (GBIF)	No associated record in PBDB
Beckwithiidae family	accepted (GBIF)	No associated record in PBDB
Bomakellidae family	accepted (GBIF)	No associated record in PBDB
Bourletiellida family	accepted (GBIF)	No associated record in PBDB
Burgessiidae family	accepted (GBIF)	No associated record in PBDB
Cetiocaridae family	accepted (GBIF)	No associated record in PBDB
Chengjiangocarididae family	accepted (GBIF)	No associated record in PBDB
Concavicarididae family	accepted (GBIF)	No associated record in PBDB
Corcoraniidae family	accepted (GBIF)	No associated record in PBDB
Cowiedesmidae family	accepted (GBIF)	No associated record in PBDB
Dicyrtomida family	accepted (GBIF)	No associated record in PBDB
Enaliktidae family	accepted (GBIF)	No associated record in PBDB
Eoarthropleuridae family	accepted (GBIF)	No associated record in PBDB
Euphoberidae family	accepted (GBIF)	No associated record in PBDB
Euthycarcinidae family	accepted (GBIF)	No associated record in PBDB
Fortiforcipidae family	accepted (GBIF)	No associated record in PBDB
Fuxianhuiidae family	accepted (GBIF)	No associated record in PBDB
Guangweicarididae ✝ family	accepted (GBIF) listed (PBDB)		1	516 Ma Cambrian	516 Ma Cambrian
Hurdiidae family	accepted (GBIF)	No associated record in PBDB
Katiannida family	accepted (GBIF)	No associated record in PBDB
Kootenichelidae family	accepted (GBIF)	No associated record in PBDB
Kottixerxidae family	accepted (GBIF)	No associated record in PBDB
Leanchoiliidae family	accepted (GBIF)	No associated record in PBDB
Marrellidae family	accepted (GBIF)	No associated record in PBDB
Microcarididae family	accepted (GBIF)	No associated record in PBDB
Microdecemplicidae family	accepted (GBIF)	No associated record in PBDB
Mictomeridae family	accepted (GBIF)	No associated record in PBDB
Neelida family	accepted (GBIF)	No associated record in PBDB
Oelandocarididae family	accepted (GBIF)	No associated record in PBDB
Offacolidae family	accepted (GBIF)	No associated record in PBDB
Onychopterellidae family	accepted (GBIF)	No associated record in PBDB
Opabiniidae family	accepted (GBIF)	No associated record in PBDB
Ostenocarididae family	accepted (GBIF)	No associated record in PBDB
Paleomeridae family	accepted (GBIF)	No associated record in PBDB
Parvancorinidae family	accepted (GBIF)	No associated record in PBDB
Pleurojulidae family	accepted (GBIF)	No associated record in PBDB
Praentombryidae family	accepted (GBIF)	No associated record in PBDB
Proiulidae family	accepted (GBIF)	No associated record in PBDB
Proscorpionidae family	accepted (GBIF)	No associated record in PBDB
Rogerellidae family	accepted (GBIF)	No associated record in PBDB
Sidneyiidae family	accepted (GBIF)	No associated record in PBDB
Sinaglaspidae family	accepted (GBIF)	No associated record in PBDB
Slimonidae family	accepted (GBIF)	No associated record in PBDB
Sminthurida family	accepted (GBIF)	No associated record in PBDB
Sminthuridida family	accepted (GBIF)	No associated record in PBDB
Sottyxerxidae family	accepted (GBIF)	No associated record in PBDB
Sprigginidae family	accepted (GBIF)	No associated record in PBDB
Strabopidae family	accepted (GBIF)	No associated record in PBDB
Waptiidae family	accepted (GBIF)	No associated record in PBDB
Yohoiidae family	accepted (GBIF)	No associated record in PBDB
Actaeus ✝ genus	accepted (GBIF) listed (PBDB)
Actoecia genus	accepted (GBIF)	No associated record in PBDB
Alaskalethe genus	accepted (GBIF)	No associated record in PBDB
Albadesmus genus	accepted (GBIF)	No associated record in PBDB
Almenia ✝ genus	accepted (GBIF) listed (PBDB)		1	520 Ma Cambrian	520 Ma Cambrian
Americotrimeria genus	accepted (GBIF)	No associated record in PBDB
Anatifera genus	accepted (GBIF)	No associated record in PBDB
Anatium genus	accepted (GBIF)	No associated record in PBDB
Anthracodesmus genus	accepted (GBIF)	No associated record in PBDB
Arator genus	accepted (GBIF)	No associated record in PBDB
Archaeopilumnus genus	accepted (GBIF)	No associated record in PBDB
Archeobius genus	accepted (GBIF)	No associated record in PBDB
Archicambala genus	accepted (GBIF)	No associated record in PBDB
Aristocaris ✝ genus	listed (PBDB)
Arlea genus	accepted (GBIF)	No associated record in PBDB
Auricula genus	accepted (GBIF)	No associated record in PBDB
Australimicola genus	accepted (GBIF)	No associated record in PBDB
Awaria genus	accepted (GBIF)	No associated record in PBDB
Baltocyamus genus	accepted (GBIF)	No associated record in PBDB
Bidentia genus	accepted (GBIF)	No associated record in PBDB
Blodgettia genus	accepted (GBIF)	No associated record in PBDB
Bolina genus	accepted (GBIF)	No associated record in PBDB
Bostrichopus ✝ genus	listed (PBDB)
Bundenbachiellus ✝ genus	listed (PBDB)
Burgessia ✝ genus	accepted (GBIF) listed (PBDB)		24	513 Ma Cambrian	513 Ma Cambrian
Calycoscutellum genus	accepted (GBIF)	No associated record in PBDB
Cambropachycope ✝ genus	listed (PBDB)
Camptocladus genus	accepted (GBIF)	No associated record in PBDB
Capayanichnus genus	accepted (GBIF)	No associated record in PBDB
Carbotubulus genus	accepted (GBIF)	No associated record in PBDB
Carpiliopsis genus	accepted (GBIF)	No associated record in PBDB
Caryosyntrips genus	accepted (GBIF)	No associated record in PBDB
Cephaloculus genus	accepted (GBIF)	No associated record in PBDB
Cephaloxus genus	accepted (GBIF)	No associated record in PBDB
Chagosia genus	accepted (GBIF)	No associated record in PBDB
Chlupaciparia genus	accepted (GBIF)	No associated record in PBDB
Chraspedops genus	accepted (GBIF)	No associated record in PBDB
Cindarella ✝ genus	accepted (GBIF) listed (PBDB)		11	520 Ma Cambrian	516 Ma Cambrian
Cineras genus	accepted (GBIF)	No associated record in PBDB
Clausocaris genus	accepted (GBIF)	No associated record in PBDB
Clypecaris ✝ genus	listed (PBDB)
Concavicaris genus	accepted (GBIF)	No associated record in PBDB
Conchotrya genus	accepted (GBIF)	No associated record in PBDB
Conia genus	accepted (GBIF)	No associated record in PBDB
Convexicaris genus	accepted (GBIF)	No associated record in PBDB
Corcorania ✝ genus	listed (PBDB)
Cornia genus	accepted (GBIF)	No associated record in PBDB
Corobalanus genus	accepted (GBIF)	No associated record in PBDB
Cretoperipatus genus	accepted (GBIF)	No associated record in PBDB
Cryptocaris ✝ genus	listed (PBDB)
Cryptozoe genus	accepted (GBIF)	No associated record in PBDB
Cultrops genus	accepted (GBIF)	No associated record in PBDB
Cyclestherioides genus	accepted (GBIF)	No associated record in PBDB
Dagoerayella genus	accepted (GBIF)	No associated record in PBDB
Dalmaniopsis ✝ genus	listed (PBDB)
Dicerocaris ✝ genus	accepted (GBIF) listed (PBDB)
Dicranocaris genus	accepted (GBIF)	No associated record in PBDB
Dicranocentroides genus	accepted (GBIF)	No associated record in PBDB
Dioxycaris genus	accepted (GBIF)	No associated record in PBDB
Donaldsoncythere genus	accepted (GBIF)	No associated record in PBDB
Dongshanocaris ✝ genus	accepted (GBIF) listed (PBDB)		1	516 Ma Cambrian	516 Ma Cambrian
Donovania genus	accepted (GBIF)	No associated record in PBDB
Douglasocaris ✝ genus	listed (PBDB)		1	468 Ma Middle Ordovician	468 Ma Middle Ordovician
Dunopyge genus	accepted (GBIF)	No associated record in PBDB
Duslia ✝ genus	listed (PBDB)		1	461 Ma Middle Ordovician	461 Ma Middle Ordovician
Editella genus	accepted (GBIF)	No associated record in PBDB
Eileticus genus	accepted (GBIF)	No associated record in PBDB
Emeiella ✝ genus	listed (PBDB)
Emeraldoides genus	accepted (GBIF)	No associated record in PBDB
Eobalanus genus	accepted (GBIF)	No associated record in PBDB
Eojulus genus	accepted (GBIF)	No associated record in PBDB
Ercaia ✝ genus	listed (PBDB)		2	520 Ma Cambrian	520 Ma Cambrian
Esmeraldacaris ✝ genus	listed (PBDB)		3	485 Ma Cambrian	485 Ma Cambrian
Estheriellites genus	accepted (GBIF)	No associated record in PBDB
Estoniosylthere genus	accepted (GBIF)	No associated record in PBDB
Euacanthus genus	accepted (GBIF)	No associated record in PBDB
Eulimene genus	accepted (GBIF)	No associated record in PBDB
Eumorphia genus	accepted (GBIF)	No associated record in PBDB
Feistia genus	accepted (GBIF)	No associated record in PBDB
Folguerolesia genus	accepted (GBIF)	No associated record in PBDB
Forfexicaris ✝ genus	listed (PBDB)		1	513 Ma Cambrian	513 Ma Cambrian
Gammarus genus	listed (PBDB)				0 Ma Extant
Gaoqiaioella ✝ genus	listed (PBDB)
Gatuna genus	accepted (GBIF)	No associated record in PBDB
Glypheopsis genus	accepted (GBIF)	No associated record in PBDB
Goticaris ✝ genus	listed (PBDB)
Gudralisium genus	accepted (GBIF)	No associated record in PBDB
Gymnolepas genus	accepted (GBIF)	No associated record in PBDB
Habelia ✝ genus	accepted (GBIF) listed (PBDB)		14	513 Ma Cambrian	513 Ma Cambrian
Hadranax ✝ genus	listed (PBDB)		2	530 Ma Cambrian	520 Ma Cambrian
Haikoucaris genus	accepted (GBIF)	No associated record in PBDB
Hanburia ✝ genus	listed (PBDB)		4	513 Ma Cambrian	513 Ma Cambrian
Hastimima genus	accepted (GBIF)	No associated record in PBDB
Helmetia genus	accepted (GBIF)	No associated record in PBDB
Hemicycloleaia genus	accepted (GBIF)	No associated record in PBDB
Hemilithobius genus	accepted (GBIF)	No associated record in PBDB
Hemiphoberia genus	accepted (GBIF)	No associated record in PBDB
Henningsmoenicaris ✝ genus	listed (PBDB)
Heterovorhoeffia genus	accepted (GBIF)	No associated record in PBDB
Houghtonites genus	accepted (GBIF)	No associated record in PBDB
Huangshandongia ✝ genus	listed (PBDB)
Ilyodes genus	accepted (GBIF)	No associated record in PBDB
Isoxys ✝ genus	accepted (GBIF) listed (PBDB)		102	530 Ma Cambrian	513 Ma Cambrian
Jianfengia ✝ genus	accepted (GBIF) listed (PBDB)		5	520 Ma Cambrian	516 Ma Cambrian
Jianshanopodia ✝ genus	listed (PBDB)
Jiucunella ✝ genus	listed (PBDB)		21	530 Ma Cambrian	520 Ma Cambrian
Juloites genus	accepted (GBIF)	No associated record in PBDB
Kablikia ✝ genus	listed (PBDB)
Kanabius genus	accepted (GBIF)	No associated record in PBDB
Kerygmachela genus	accepted (GBIF)	No associated record in PBDB
Kirkbyrhiza genus	accepted (GBIF)	No associated record in PBDB
Kleptothule ✝ genus	accepted (GBIF) listed (PBDB)		1	520 Ma Cambrian	520 Ma Cambrian
Kralupia genus	accepted (GBIF)	No associated record in PBDB
Krauselloides genus	accepted (GBIF)	No associated record in PBDB
Kuamaia genus	accepted (GBIF)	No associated record in PBDB
Laevanotella genus	accepted (GBIF)	No associated record in PBDB
Leptophilus genus	accepted (GBIF)	No associated record in PBDB
Leverhulmia genus	accepted (GBIF)	No associated record in PBDB
Liantuoia ✝ genus	listed (PBDB)
Lingyuanaspis genus	accepted (GBIF)	No associated record in PBDB
Lockingtonia genus	accepted (GBIF)	No associated record in PBDB
Longquania ✝ genus	accepted (GBIF) listed (PBDB)		1	516 Ma Cambrian	516 Ma Cambrian
Loricicaris genus	accepted (GBIF)	No associated record in PBDB
Luohuilinella genus	accepted (GBIF)	No associated record in PBDB
Machairopus genus	accepted (GBIF)	No associated record in PBDB
Maldybulakia genus	accepted (GBIF)	No associated record in PBDB
Marrella ✝ genus	listed (PBDB)		29	513 Ma Cambrian	513 Ma Cambrian
Marria genus	accepted (GBIF)	No associated record in PBDB
Martinssonia ✝ genus	listed (PBDB)		1	501 Ma Cambrian	501 Ma Cambrian
Mayrocaris genus	accepted (GBIF)	No associated record in PBDB
Megadictyon ✝ genus	accepted (GBIF) listed (PBDB)		2	520 Ma Cambrian	520 Ma Cambrian
Meristosomo ✝ genus	listed (PBDB)
Mesoporcellio genus	accepted (GBIF)	No associated record in PBDB
Mesosoma genus	accepted (GBIF)	No associated record in PBDB
Molaria ✝ genus	listed (PBDB)		19	513 Ma Cambrian	513 Ma Cambrian
Mollisonia ✝ genus	listed (PBDB)		7	513 Ma Cambrian	513 Ma Cambrian
Monoleiolophus genus	accepted (GBIF)	No associated record in PBDB
Montcestheria genus	accepted (GBIF)	No associated record in PBDB
Motina ✝ genus	listed (PBDB)
Naptile ✝ genus	listed (PBDB)
Necrozius genus	accepted (GBIF)	No associated record in PBDB
Nectotelson genus	accepted (GBIF)	No associated record in PBDB
Nelocira genus	accepted (GBIF)	No associated record in PBDB
Neocypris genus	accepted (GBIF)	No associated record in PBDB
Nereocaris genus	accepted (GBIF)	No associated record in PBDB
Nettapezoura genus	accepted (GBIF)	No associated record in PBDB
Notopella genus	accepted (GBIF)	No associated record in PBDB
Occacaris ✝ genus	listed (PBDB)
Odaraia ✝ genus	accepted (GBIF) listed (PBDB)		18	520 Ma Cambrian	513 Ma Cambrian
Odonterichthus genus	accepted (GBIF)	No associated record in PBDB
Oelandocaris genus	accepted (GBIF)	No associated record in PBDB
Onebius genus	accepted (GBIF)	No associated record in PBDB
Oniscina genus	accepted (GBIF)	No associated record in PBDB
Onychocythere genus	accepted (GBIF)	No associated record in PBDB
Oodeopus genus	accepted (GBIF)	No associated record in PBDB
Otion genus	accepted (GBIF)	No associated record in PBDB
Oxyuropoda ✝ genus	listed (PBDB)
Pagurus genus	listed (PBDB)		42	161 Ma Jurassic	0 Ma Extant
Pahvantia ✝ genus	accepted (GBIF) listed (PBDB)
Palaeodesmus genus	accepted (GBIF)	No associated record in PBDB
Pamina genus	accepted (GBIF)	No associated record in PBDB
Panlongia ✝ genus	accepted (GBIF) listed (PBDB)		3	516 Ma Cambrian	516 Ma Cambrian
Paracreusia genus	accepted (GBIF)	No associated record in PBDB
Paraostenia genus	accepted (GBIF)	No associated record in PBDB
Parapaleomerus ✝ genus	accepted (GBIF) listed (PBDB)		3	520 Ma Cambrian	516 Ma Cambrian
Parathanas genus	accepted (GBIF)	No associated record in PBDB
Parodolepas genus	accepted (GBIF)	No associated record in PBDB
Parvancorina genus	accepted (GBIF)	No associated record in PBDB
Paulinecaris genus	accepted (GBIF)	No associated record in PBDB
Pauloterminus genus	accepted (GBIF) listed (PBDB)		2	530 Ma Cambrian	520 Ma Cambrian
Pectocaris ✝ genus	accepted (GBIF) listed (PBDB)		6	541 Ma Cambrian	516 Ma Cambrian
Pemphicyclus genus	accepted (GBIF)	No associated record in PBDB
Peteinura genus	accepted (GBIF)	No associated record in PBDB
Phylluropus genus	accepted (GBIF)	No associated record in PBDB
Phytophilaspis genus	accepted (GBIF)	No associated record in PBDB
Pisinnocaris ✝ genus	listed (PBDB)		2	541 Ma Cambrian	541 Ma Cambrian
Plagiolophus genus	accepted (GBIF)	No associated record in PBDB
Platicytes genus	accepted (GBIF) listed (PBDB)		1	359 Ma Late/Upper Devonian	359 Ma Late/Upper Devonian
Plenocaris ✝ genus	accepted (GBIF) listed (PBDB)		16	513 Ma Cambrian	513 Ma Cambrian
Pleuroschendyla genus	accepted (GBIF)	No associated record in PBDB
Pogonoligia genus	accepted (GBIF)	No associated record in PBDB
Polzia genus	accepted (GBIF)	No associated record in PBDB
Praelepas genus	accepted (GBIF)	No associated record in PBDB
Priscansermarinus ✝ genus	listed (PBDB)		2	513 Ma Cambrian	513 Ma Cambrian
Proboliella genus	accepted (GBIF)	No associated record in PBDB
Propemesira genus	accepted (GBIF)	No associated record in PBDB
Protocarcinus genus	listed (PBDB)		1	170 Ma Middle Jurassic	170 Ma Middle Jurassic
Protocaris ✝ genus	listed (PBDB)		3	530 Ma Cambrian	516 Ma Cambrian
Protozoe genus	accepted (GBIF)	No associated record in PBDB
Protozoea genus	accepted (GBIF)	No associated record in PBDB
Pseudarthron ✝ genus	listed (PBDB)
Pseuderichthus genus	accepted (GBIF)	No associated record in PBDB
Pseudoarctolepis ✝ genus	listed (PBDB)		2	513 Ma Cambrian	513 Ma Cambrian
Pseudoasmussia genus	accepted (GBIF)	No associated record in PBDB
Pseudocypridina genus	accepted (GBIF)	No associated record in PBDB
Pseudoiulia ✝ genus	accepted (GBIF) listed (PBDB)		4	516 Ma Cambrian	516 Ma Cambrian
Pseudontella genus	accepted (GBIF)	No associated record in PBDB
Pseudovendia genus	accepted (GBIF)	No associated record in PBDB
Pterisopodus genus	accepted (GBIF)	No associated record in PBDB
Pugilator genus	accepted (GBIF)	No associated record in PBDB
Pygmaclypeatus ✝ genus	listed (PBDB)
Quasicaris ✝ genus	listed (PBDB)		1	461 Ma Middle Ordovician	461 Ma Middle Ordovician
Raninellopsis genus	accepted (GBIF)	No associated record in PBDB
Rehbachiella ✝ genus	listed (PBDB)		1	501 Ma Cambrian	501 Ma Cambrian
Retifacies ✝ genus	accepted (GBIF) listed (PBDB)		18	520 Ma Cambrian	516 Ma Cambrian
Retiprimites genus	accepted (GBIF)	No associated record in PBDB
Rhinediaptomus genus	accepted (GBIF)	No associated record in PBDB
Rhombicalvaria ✝ genus	accepted (GBIF) listed (PBDB)		2	520 Ma Cambrian	520 Ma Cambrian
Rhynimonstrum ✝ genus	accepted (GBIF) listed (PBDB)		1	411 Ma Early/Lower Devonian	411 Ma Early/Lower Devonian
Roddyia genus	accepted (GBIF)	No associated record in PBDB
Sagittapeltis genus	accepted (GBIF)	No associated record in PBDB
Sandtneria genus	accepted (GBIF)	No associated record in PBDB
Schinderhannes ✝ genus	listed (PBDB)		1	408 Ma Early/Lower Devonian	408 Ma Early/Lower Devonian
Schizodiscus ✝ genus	accepted (GBIF) listed (PBDB)		3	388 Ma Middle Devonian	388 Ma Middle Devonian
Serracaris ✝ genus	listed (PBDB)		1	516 Ma Cambrian	516 Ma Cambrian
Shafferia ✝ genus	accepted (GBIF) listed (PBDB)
Shankouia ✝ genus	accepted (GBIF) listed (PBDB)		3	520 Ma Cambrian	516 Ma Cambrian
Siberion ✝ genus	listed (PBDB)
Sinoburius ✝ genus	accepted (GBIF) listed (PBDB)		3	520 Ma Cambrian	520 Ma Cambrian
Siphonocetes genus	accepted (GBIF)	No associated record in PBDB
Skioldia genus	accepted (GBIF)	No associated record in PBDB
Smithixerxes genus	accepted (GBIF)	No associated record in PBDB
Sottyxerxes genus	accepted (GBIF)	No associated record in PBDB
Spathiocaris genus	accepted (GBIF)	No associated record in PBDB
Spencebatea genus	accepted (GBIF)	No associated record in PBDB
Spinifer genus	accepted (GBIF)	No associated record in PBDB
Squama genus	accepted (GBIF)	No associated record in PBDB
Squamacula genus	accepted (GBIF)	No associated record in PBDB
Strongylonotus genus	accepted (GBIF)	No associated record in PBDB
Struveops genus	accepted (GBIF)	No associated record in PBDB
Suchonella genus	accepted (GBIF)	No associated record in PBDB
Tanglangia ✝ genus	listed (PBDB)		2	520 Ma Cambrian	516 Ma Cambrian
Tetrathrosoma genus	accepted (GBIF)	No associated record in PBDB
Thaliella genus	accepted (GBIF)	No associated record in PBDB
Thylacares genus	accepted (GBIF)	No associated record in PBDB
Thylacocephalus genus	accepted (GBIF)	No associated record in PBDB
Trichelaspis genus	accepted (GBIF)	No associated record in PBDB
Trichiulus genus	accepted (GBIF)	No associated record in PBDB
Trichorypha genus	accepted (GBIF)	No associated record in PBDB
Urokodia ✝ genus	accepted (GBIF) listed (PBDB)		6	520 Ma Cambrian	516 Ma Cambrian
Utahcaris ✝ genus	accepted (GBIF) listed (PBDB)		1	513 Ma Cambrian	513 Ma Cambrian
Vachonisia genus	accepted (GBIF)	No associated record in PBDB
Victoriacaris genus	accepted (GBIF)	No associated record in PBDB
Walossekia ✝ genus	listed (PBDB)		1	501 Ma Cambrian	501 Ma Cambrian
Weyerites genus	accepted (GBIF)	No associated record in PBDB
Whipplella genus	accepted (GBIF)	No associated record in PBDB
Wiedopterus genus	accepted (GBIF)	No associated record in PBDB
Xandarella ✝ genus	accepted (GBIF) listed (PBDB)		15	520 Ma Cambrian	516 Ma Cambrian
Zoontocaris genus	accepted (GBIF)	No associated record in PBDB
Actinobalnus collinsi species	accepted (GBIF)	No associated record in PBDB
Actinobalnus floridanus species	accepted (GBIF)	No associated record in PBDB
Alalcomenaeus cambricus species	accepted (GBIF)	No associated record in PBDB
Cestites mirebilis species	accepted (GBIF)	No associated record in PBDB
Cylestherioides pintoi species	accepted (GBIF)	No associated record in PBDB
Enosiaspis hrungnir species	accepted (GBIF)	No associated record in PBDB
Entocaris telleri species	accepted (GBIF)	No associated record in PBDB
Euscaplellum wheeleri species	accepted (GBIF)	No associated record in PBDB
Huxleyoecia muscatensis species	accepted (GBIF)	No associated record in PBDB
Hyperbionx pluto species	accepted (GBIF)	No associated record in PBDB
Idiotheca rugosa species	accepted (GBIF)	No associated record in PBDB
Kouphichnium arizonae species	accepted (GBIF)	No associated record in PBDB
Lisgocaris lutheri species	accepted (GBIF)	No associated record in PBDB
Pachycrommium princetonensis species	accepted (GBIF)	No associated record in PBDB
Perspicaris dictynna species	accepted (GBIF)	No associated record in PBDB
Perspicaris dilatus species	accepted (GBIF)	No associated record in PBDB
Perspicaris ellipsopelta species	accepted (GBIF)	No associated record in PBDB
Perspicaris recondita species	accepted (GBIF)	No associated record in PBDB
Platylepis wilsoni species	accepted (GBIF)	No associated record in PBDB
Redondestheria grovetonensis species	accepted (GBIF)	No associated record in PBDB
Sarotrocercus oblita species	accepted (GBIF)	No associated record in PBDB
Stetaspis spinulosus species	accepted (GBIF)	No associated record in PBDB
Sudanonauter kagoroensis species	accepted (GBIF)	No associated record in PBDB
Branchiata unranked clade	listed (PBDB)				0 Ma Extant
Conchostraca unranked clade	listed (PBDB)	Clam Shrimp	71	541 Ma Cambrian	0 Ma Extant
Parvancorinomorpha ✝ unranked clade	listed (PBDB)		20	520 Ma Cambrian	513 Ma Cambrian

Synonymy List

Year	Name and Author
1758	Aptera Linnaeus p. 341
1758	Monoculus Linnaeus p. 634
1761	Aptera Linnaeus p. 472
1761	Monoculus Linnaeus p. 497
1762	Monoculus Geoffroy p. 651
1763	Pedestria Scopoli p. 378
1763	Monoculus Scopoli p. 413
1764	Aptera Müller p. 89
1764	Monoculus Müller p. 95
1766	Aptera Schaeffer
1767	Aptera Linnaeus p. 1012
1767	Monoculus Linnaeus p. 1057
1768	Aptera Strøm p. 359
1770	Aptera Drury
1775	Monoculus Fabricius p. 294
1775	Synistata Fabricius p. 294
1775	Agonata Fabricius p. 399
1775	Unogata Fabricius p. 420
1776	Aptera Müller p. 183
1798	Synistata Fabricius p. 197
1829	Arthropoda Latreille

Year	Name and Author
1865	Blattina splendens Göppert p. 276 figs. Pl 64, fig 11
1876	Libellula carbonaria Scudder p. 89 fig. 2
1890	Graeophonus Scudder p. 453
1890	Graeophonus carbonarius Scudder p. 454 figs. Pl 40, figs 2,3,6
1897	Geracus Matthew p. 55
1897	Geracus tubifer Matthew p. 55 fig. 3
1899	Kampecaris obanensis Peach p. 122 figs. Pl 4, fig 2
1901	Kounicia Fric p. 175
1913	Graeophonus carbonarius Petrunkevitch p. 67 figs. Pl 5, figs 27-29; text-figs 29-31
1953	Graeophonus carbonarius Petrunkevitch p. 99
1963	Poliochera pustulatus Laurentiaux-Vieira and Laurentiaux p. 25 figs. Pl 3, fig 3; text-fig 3
1976	Schizoramia Bergström
1989	Arthropoda Shoshani et al. p. 435
1998	Arthropoda Zrzavy et al.
2002	Arthropoda Sepkoski, Jr.
2011	Arthropoda Edgecombe et al. p. 153 figs. Fig. 1
2012	Arthropoda Nielsen p. 240 figs. Fig. 42.1
2013	Graeophonus Dunlop et al. p. 190
2013	Graeophonus carbonarius Dunlop et al. p. 190
2014	Graeophonus Engel and Grimaldi p. 3
2017	Graeophonus Garwood et al. p. 12

References

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Garwood R. J., Dunlop J. A., et al (2017) The phylogeny of fossil whip spiders, BMC Evolutionary Biology 17 105, 1-14

Engel M. S., Grimaldi D. A. (2014) Whipspiders (Arachnida: Amblypygi) in amber from the early Eocene and mid-Cretaceous, including maternal care, Novitates Paleoentomologicae 9, 1-17

Dunlop J. A., Penney D., et al (2013) A summary list of fossil spiders and their relatives, The World Spider Catalog, version 14.0. American Museum of Natural History

Nielsen C. (2012) Animal Evolution : Interrelationships of the Living Phyla, 1-402

Edgecombe G. D., Giribet G., et al (2011) Higher-level metazoan relationships: recent progress and remaining questions, Organisms Diversity & Evolution 11 2, 151-172 doi:10.1007/s13127-011-0044-4

Zrzavy J., Mihulka S., et al (1998) Phylogeny of the Metazoa based on morphological and 18S ribosomal DNA evidence, Cladistics 14, 249-285

Shoshani J., Fisher D. C., et al (1989) The Shelton Mastodon Site: Multidisciplinary study of a late Pleistocene (Twocreekan) locality in Southeastern Michigan, Contributions from the Museum of Paleontology, University of Michigan 27 14, 393-436

Laurentiaux-Vieira F., Laurentiaux D. (1963) Sur quelques restes noveaux d'Arachnides du terrain houiller, Annales de la Société Géologique du Nord 83, 23-29

Petrunkevitch A. I. (1953) Paleozoic and Mesozoic Arachnida of Europe, Geological Society of America Memoir 53, 1-122

Petrunkevitch A. I. (1913) A monograph of the terrestrial Palaeozoic Arachnida of North America, Transactions of the Connecticut Acadamy of Arts and Sciences 18, 1-137

Fric A. (1901) Die thierischen Reste der Perucer Schichten, Archiv für die Naturwissenschaftliche Landesdurchforschung von Böhmen 11, 163-180

Peach B. N. On some new myriapods from the Palaeozoic rocks of Scotland, Proceedings of the Royal Physical Society of Edinburgh 14, 113-126

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Linnaeus C. , Fauna Svecica: Sistens Animalia Sveciae Regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes, Distributa per Classes & Ordines, Genera & Species, cum Differentiis Specierum, Synonymis Auctorum, Nominibus Incolarum, Locis Natalium, Descriptionibus Insectorum, 1-578

Linnaeus C. , Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1, 1-824 doi:10.5962/bhl.title.542

Ringuelet,Raul A. (1947) Aceros e insectos parásitos, nuevos para la fauna argentina - via Phthiraptera.info

Pavlovic,I.,Kulišić,Z.,Milutinovi,Ć.,Nešić,D. :. Valter,D. (1995) Significance of arthropoda in health problems of pheasants which are bred artificially - via Phthiraptera.info

Hayward, P.J.; Ryland, J.S. (Ed.). (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. Clarendon Press: Oxford, UK. ISBN 0-19-857356-1. 627 pp. - via World Register of Introduced Marine Species (WRiMS)

Margulis, L.; Schwartz, K.V. (1998). Five Kingdoms: an illustrated guide to the Phyla of life on earth. 3rd edition. Freeman: New York, NY (USA). ISBN 0-7167-3027-8. xx, 520 pp. - via World Register of Introduced Marine Species (WRiMS)

Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas

Eichler,Wolfdietrich,Bronns,H. G. (1963) Arthropoda. Insecta. Phthiraptera I. Mallophaga. - via Phthiraptera.info

Cloudsley-Thompson,J. L. (1975) Adaptations of Arthropoda to arid environments - via Phthiraptera.info

Urbanski,F. (1979) Arthropoda collected on small mammals in Wielkopolski. National Parc - via Phthiraptera.info

Baccetti,Baccio,Dallai,Romano,Rosati,Floriana (1969) The spermatozoon of Arthropoda. IV. Corrodentia, Mallophaga and Thysanoptera - via Phthiraptera.info

Beutel,Rolf G.,Gorb,Stanislav N. (2001) Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny - via Phthiraptera.info

Kim,Ke Chung,Beaucournu,Jean-Claude,Doby,J. M.,Kremer,M.,Leger,N.,Leclercq,M.,Olufsjev,N. G.,Challier,A.,Fain,A. (1978) Morphology and taxonomy of parasitic arthropoda - via Phthiraptera.info

Ricardo L. Palma,Lovis,P. M.,Tither,C. (1989) An annotated list of primary types of the phyla Arthropoda (except Crustacea) and Tardigrada held in the National Museum of New Zealand - via Phthiraptera.info

Olmos,Viviana,Muñoz,Gabriela (2006) Current state of knowledge of eumetazoan parasites of chilean freshwater ecosystems - via Phthiraptera.info

Dimitri Ivanovich Blagoveshtchensky (1937) Materialy po faune naruznych parasitov (Arthropoda) zyvotnych kasalinskogo i nekotorych drugich rajonov juznogo Kasakstana. Materialy po vreditelam zyvotnovodstva i fauna preimuscestvenno juznogo Kasakstana, pod red - via Phthiraptera.info

Green,R. H.,Munday,B. L. (1971) Parasites of Tasmanian native and feral fauna. Part 1. Arthropoda - via Phthiraptera.info

Hansen,Hans Jakob (1930) Studies on Arthropoda. III. On the comparative morphology of the appendages of the Arthropoda. B. Crustacea, Insecta, Myriopoda, and Arachnida - via Phthiraptera.info

Brusca, Richard C., and Gary J. Brusca, 1990: null. Invertebrates. xiii + 992. - via Integrated Taxonomic Information System (ITIS)

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Stephen C. Barker,Campbell,N. J. H. (1998) An unprecedented major rearrangement in an arthropod mitochondrial genome - via Phthiraptera.info

Złotorzycka,Jadwiga (1972) Problemy wspolzaleznosci ewolucji Arthropoda i ich zywicieli - via Phthiraptera.info

Hegna, T. A.; Legg, D. A.; Møller, O. S.; Van Roy, P.; Lerosey-Aubril, R. (2013). The correct authorship of the taxon name 'Arthropoda'. Arthropod Systematics & Philogeny. 71(2), 71-74. - via World Register of Introduced Marine Species (WRiMS)

Zhang, Z.-Q. (2013). Phylum Arthropoda. In: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa. 3703(1): 17-26. - via The Interim Register of Marine and Nonmarine Genera

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Ruggiero, M. A.; Gordon, D. P.; Orrell, T. M.; Bailly, N.; Bourgoin, T.; Brusca, R. C.; Cavalier-Smith, T.; Guiry, M. D.; Kirk, P. M. (2015). A higher level classification of all living organisms. PLOS ONE. 10(4): e0119248. - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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