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Heteroceridae

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Description

The Heteroceridae, or variegated mud-loving beetles, are a widespread and relatively common family of beetles. They occur on every continent except for Antarctica.

From Wikipedia article at https://en.wikipedia.org/wiki/Heteroceridae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	6986	https://www.gbif.org/species/6986
PaleoBioDB ID (PBDB)	69343	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=69343

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Heteroceridae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Polyphaga	Emery 1886
infraorder	Elateriformia	Crowson 1960
superfamily	Byrrhoidea	Latreille 1804
family	Heteroceridae	MacLeay 1825

Common Name

Variegated mud-loving beetle

Scientific Name

Heteroceridae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Heteroceridae	family	belongs to Dryopoidea	stated without evidence	Crowson, 1981
Heteroceridae	family	belongs to Byrrhoidea	stated without evidence	Lawrence and Newton, 1995
Heteroceridae	family	belongs to Byrrhoidea	stated with evidence	Bouchard et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

12 (80%)

First Recorded Appearance

125 - 122 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

R. A. Crowson. 1981. The Biology of the Coleoptera. Academic Press, New York

Common Name(s)

Oevergraafkevers, Variegated Mud-loving Beetles, Variegated Mud-loving Beetle, Strandgrävbaggar, Mud Beetles, Strandgravarar, Strandgravere

Wikipedia

https://en.wikipedia.org/wiki/Heteroceridae

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Elythomerinae subfamily	listed (PBDB)				0 Ma Extant
Heterocerinae subfamily	listed (PBDB)		8	0.78 Ma Pleistocene	0 Ma Extant
Augyles genus	accepted (GBIF)	No associated record in PBDB
Centuriatus genus	accepted (GBIF)	No associated record in PBDB
Efflagitatus genus	accepted (GBIF)	No associated record in PBDB
Explorator genus	accepted (GBIF)	No associated record in PBDB
Heterocerites ✝ genus	accepted (GBIF) listed (PBDB)		2	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Heterocerus genus	accepted (GBIF) listed (PBDB)		8	0.78 Ma Pleistocene	0 Ma Extant
Lanternarius genus	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Lapsus genus	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Littorimus genus	accepted (GBIF)	No associated record in PBDB
Micilus genus	accepted (GBIF)	No associated record in PBDB
Microaugyles genus	accepted (GBIF)	No associated record in PBDB
Neoheterocerus genus	accepted (GBIF)	No associated record in PBDB
Peditatus genus	accepted (GBIF)	No associated record in PBDB
Protoallopygmephorus genus	accepted (GBIF)	No associated record in PBDB
Tropicus genus	accepted (GBIF)	No associated record in PBDB
Culnus sinaloensis species	accepted (GBIF)	No associated record in PBDB

Synonymy List

Year	Name and Author
1825	Heteroceridae MacLeay
1981	Heteroceridae Crowson p. 695
1995	Heteroceridae Lawrence and Newton, Jr.
2011	Heteroceridae Bouchard et al. p. 49

References

Crowson R. A. (1981) The Biology of the Coleoptera, Academic Press, New York

Lawrence J. F., Newton, Jr. A. F. (1995) Families and subfamilies of Coleoptera, Biology, Phylogeny and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A Crowson, 779-1006

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas

Lawrence, J. F., and A. F. Newton, Jr. / Pakaluk, James, and Stanislaw Adam Slipinski, eds., 1995: Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson, vol. 2. 779-1006. - via Integrated Taxonomic Information System (ITIS)

Katovich, Kerry / Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley, and J. H. Frank, eds., 2002: Family 47: Heteroceridae MacLeay 1825. American Beetles: Vol. 2: Polyphaga: Scarabaeoidea through Curculionoidea. 127-132. - via Integrated Taxonomic Information System (ITIS)

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera

Hayward, P.J.; Ryland, J.S. (Ed.). (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. Clarendon Press: Oxford, UK. ISBN 0-19-857356-1. 627 pp. - via World Register of Marine Species

Freude, et al. (1965) - via Taxa Watermanagement the Netherlands (TWN)

Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Slipinski, A.; Smith, A. (2011). Family-group names in Coleoptera (Insecta). ZooKeys. 88: 1-972. - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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