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Harpes ✝

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Harpes perradiatus

Morocco

Harpes perradiatus

Morocco

Harpes perradiatus

Morocco

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3266942	https://www.gbif.org/species/3266942
PaleoBioDB ID (PBDB)	20537	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=20537

Rank

genus

Taxonomy (GBIF)

Life : Animalia : Arthropoda : Trilobita : Harpetida : Harpetidae : Harpes

Taxonomy (PBDB)

Life : Animalia : Arthropoda : Trilobita : Harpida : Harpetidae : Harpes

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
class	Trilobita	Walch 1771
order	Harpida
family	Harpetidae	Hawle and Corda 1847
genus	Harpes	Goldfuss 1839

Scientific Name

Harpes Goldfuss, 1839

Name Published In

Nova Acta Academiae Caesarea Leopoldino-Carolinae Germanicum Naturae Curiosorum, 19(1): 358

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Harpes	genus	belongs to Harpetidae	stated with evidence	Ebach and McNamara, 2002
Harpes	genus	belongs to Ptychopariida	second hand	Sepkoski, 2002
Harpes	genus	belongs to Harpetidae	stated without evidence	Jell and Adrain, 2003

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

468 - 464 Ma
Middle Ordovician

Last Recorded Appearance

380 - 376 Ma
Late/Upper Devonian

Environment

marine (based on Trilobita)

Motility

fast-moving (based on Trilobita)

Vision

limited (based on Harpetidae)

Diet

suspension feeder (based on Harpetidae)

Ontogeny

molting,addition of parts (based on Trilobita)

Taphonomy

low Mg calcite, chitin (based on Trilobita)

Primary Reference (PBDB)

R. Feist. 2002. Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta). Acta Palaeontologica Polonica 47(2):203-210

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Harpes acuminatus ✝ species	accepted (GBIF) listed (PBDB)
Harpes dvorcensis ✝ species	accepted (GBIF) listed (PBDB)		1	411 Ma Early/Lower Devonian	411 Ma Early/Lower Devonian
Harpes ellipticus species	accepted (GBIF)	No associated record in PBDB
Harpes escoti ✝ species	accepted (GBIF) listed (PBDB)
Harpes fornicatus ✝ species	accepted (GBIF) listed (PBDB)
Harpes forojuliensis ✝ species	accepted (GBIF) listed (PBDB)
Harpes gracilis ✝ species	accepted (GBIF) listed (PBDB)
Harpes koeneni ✝ species	accepted (GBIF) listed (PBDB)
Harpes latilimbatus ✝ species	accepted (GBIF) listed (PBDB)
Harpes macrocephallus ✝ species	listed (PBDB)
Harpes macrocephalus ✝ species	accepted (GBIF) listed (PBDB)		6	408 Ma Early/Lower Devonian	393 Ma Early/Lower Devonian
Harpes neogracilis ✝ species	accepted (GBIF) listed (PBDB)		2	380 Ma Late/Upper Devonian	380 Ma Late/Upper Devonian
Harpes nymageenensis ✝ species	accepted (GBIF) listed (PBDB)
Harpes nymageensis ✝ species	accepted (GBIF) listed (PBDB)
Harpes ormistoni ✝ species	accepted (GBIF) listed (PBDB)
Harpes parvulus ✝ species	accepted (GBIF) listed (PBDB)
Harpes perradiatus ✝ species	accepted (GBIF) listed (PBDB)		2	393 Ma Early/Lower Devonian	393 Ma Early/Lower Devonian
Harpes polaris ✝ species	accepted (GBIF) listed (PBDB)		1	409 Ma Early/Lower Devonian	409 Ma Early/Lower Devonian
Harpes pruniformis species	accepted (GBIF)	No associated record in PBDB
Harpes pyrenaicus ✝ species	accepted (GBIF) listed (PBDB)
Harpes radians ✝ species	accepted (GBIF) listed (PBDB)
Harpes reticulatus ✝ species	listed (PBDB)		2	419 Ma Early/Lower Devonian	409 Ma Early/Lower Devonian
Harpes rouvillei ✝ species	accepted (GBIF) listed (PBDB)
Harpes socialis ✝ species	accepted (GBIF) listed (PBDB)
Harpes speciosus species	accepted (GBIF)	No associated record in PBDB
Harpes transiens ✝ species	accepted (GBIF) listed (PBDB)		3	393 Ma Early/Lower Devonian	388 Ma Middle Devonian
Harpes whidbournei ✝ species	accepted (GBIF) listed (PBDB)
Reticuloharpes reticulatus ✝ species	accepted (GBIF) listed (PBDB)

Synonymy List

Year	Name and Author
1839	Harpes Goldfuss
1963	Reticuloharpes Vanek
1981	Helioharpes Pribyl and Vanek
2002	Harpes Ebach and McNamara
2002	Harpes Sepkoski, Jr.
2002	Helioharpes Sepkoski, Jr.
2002	Reticuloharpes Sepkoski, Jr.
2003	Harpes Jell and Adrain p. 382

References

Feist R. (2002) Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta), Acta Palaeontologica Polonica 47 2, 203-210

Ebach M. C., McNamara K. J. (2002) A systematic revision of the family Harpetidae (Trilobita), Records of the Western Australian Museum 21, 135-167

Sepkoski, Jr. J. J. (2002) A compendium of fossil marine animal genera, Bulletins of American Paleontology 363, 1-560

International Commission on Zoological Nomenclature. Opinion 1436 Harpidae Hawle & Corda, 1847 (Trilobita) and Harpidae Bronn, 1849 (Mollusca, Gastropoda): a ruling to remove the homonymy.Bulletin of Zoological Nomenclature 44, 137-138 (1987) - via Official Lists and Indexes of Names in Zoology

Museum Victoria KEmu database (Oct 2006) - via The Interim Register of Marine and Nonmarine Genera

Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

Sepkoski, J. J., Jr. (2002). A compendium of fossil marine animal genera. Bulletins of American Paleontology. 363, 1-560. - via The Interim Register of Marine and Nonmarine Genera

extinct group - via The Interim Register of Marine and Nonmarine Genera

Nova Acta Acad. Caes. Leop. Carol., 19 (1) - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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