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Moghara, Egypt

Formation	Moghara
Age:	16.9 - 16.0 Ma Miocene
Interval	MN 4
Lithology	sandstone
Number of Collections	9
Number of Occurrences	47

Recorded Sample Locations

Location	Region	Stratigraphic Name
Wadi Moghra	Matrouh Governorate, Egypt	Moghara
Wadi Faregh	Beheira Governorate, Egypt	Moghara

Recorded Fossils

Accepted Name	Type	Hierarchy	Age
Lates sp. genus	6	Animalia : Chordata : Actinopterygii : Perciformes : Latidae : *Lates*	16.9 - 15.97 Ma Miocene
Synodontis sp. genus	6	Animalia : Chordata : Actinopteri : Siluriformes : Mochokidae : *Synodontis*	16.9 - 15.97 Ma Miocene
Leptoptilos sp. genus	6	Animalia : Chordata : Aves : Ciconiiformes : Ciconiidae : *Leptoptilos*	16.9 - 15.97 Ma Miocene
Sphyrna prisca species	7	Animalia : Chordata : Elasmobranchii : Carcharhiniformes : Sphyrnidae : Sphyrna : *Sphyrna prisca*	16.9 - 15.97 Ma Miocene
Myliobatis sp. genus	6	Animalia : Chordata : Elasmobranchii : Myliobatiformes : Myliobatidae : *Myliobatis*	16.9 - 15.97 Ma Miocene
Pristis sp. genus	6	Animalia : Chordata : Elasmobranchii : Rhinopristiformes : Pristidae : *Pristis*	16.9 - 15.97 Ma Miocene
Brachyodus sp. genus	6	Animalia : Chordata : Mammalia : Artiodactyla : Anthracotheriidae : *Brachyodus*	16.9 - 15.97 Ma Miocene
Brachyodus moneyi species	7	Animalia : Chordata : Mammalia : Artiodactyla : Anthracotheriidae : Brachyodus : *Brachyodus moneyi*	16.9 - 15.97 Ma Miocene
Masritherium depereti species	7	Animalia : Chordata : Mammalia : Artiodactyla : Anthracotheriidae : Masritherium : *Masritherium depereti*	16.9 - 15.97 Ma Miocene
Prolibytherium magnieri species	7	Animalia : Chordata : Mammalia : Artiodactyla : Palaeomerycidae : Prolibytherium : *Prolibytherium magnieri*	16.9 - 15.97 Ma Miocene
Cynelos sp. genus	6	Animalia : Chordata : Mammalia : Carnivora : Amphicyonidae : *Cynelos*	16.9 - 15.97 Ma Miocene
Hyaena sp. genus	6	Animalia : Chordata : Mammalia : Carnivora : Hyaenidae : *Hyaena*	16.9 - 15.97 Ma Miocene
Herpestides aegypticus species	7	Animalia : Chordata : Mammalia : Carnivora : Hyaenidae : Herpestides : *Herpestides aegypticus*	23.03 - 15.97 Ma Miocene
Moghradictis nedjema species	7	Animalia : Chordata : Mammalia : Carnivora : Stenoplesictidae : Moghradictis : *Moghradictis nedjema*	23.03 - 15.97 Ma Miocene
Ketketictis solida species	7	Animalia : Chordata : Mammalia : Carnivora : Viverridae : Ketketictis : *Ketketictis solida*	23.03 - 15.97 Ma Miocene
Odontoceti suborder	4.3	Animalia : Chordata : Mammalia : Cetacea : Champsodelphis : Odontoceti	16.9 - 15.97 Ma Miocene
Delphinus sulcatus species	7	Animalia : Chordata : Mammalia : Cetacea : Eurhinodelphinidae : Schizodelphis : *Delphinus sulcatus*	23.03 - 15.97 Ma Miocene
Hyainailouros sp. genus	6	Animalia : Chordata : Mammalia : Hyaenodonta : Hyainailouridae : *Hyainailouros*	16.9 - 15.97 Ma Miocene
Megistotherium osteothlastes species	7	Animalia : Chordata : Mammalia : Hyaenodontidae : Megistotherium : *Megistotherium osteothlastes*	16.9 - 15.97 Ma Miocene
Buhakia moghraensis species	7	Animalia : Chordata : Mammalia : Teratodontidae : Buhakia : *Buhakia moghraensis*	16.9 - 15.97 Ma Miocene
Teratodon sp. genus	6	Animalia : Chordata : Mammalia : Hyaenodontidae : *Teratodon*	23.03 - 15.97 Ma Miocene
Aceratherium sp. genus	6	Animalia : Chordata : Mammalia : Perissodactyla : Rhinocerotidae : *Aceratherium*	16.9 - 15.97 Ma Miocene
Teleoceras snowi species	7	Animalia : Chordata : Mammalia : Perissodactyla : Rhinocerotidae : Teleoceras : *Teleoceras snowi*	16.9 - 15.97 Ma Miocene
Dryopithecus mogharensis species	7	Animalia : Chordata : Mammalia : Primates : Hominidae : Dryopithecus : *Dryopithecus mogharensis*	16.9 - 15.97 Ma Miocene
Prohylobates tandyi species	7	Animalia : Chordata : Mammalia : Primates : Cercopithecidae : Prohylobates : *Prohylobates tandyi*	16.9 - 15.97 Ma Miocene
Archaeobelodon sp. genus	6	Animalia : Chordata : Mammalia : Proboscidea : Gomphotheriidae : *Archaeobelodon*	16.9 - 15.97 Ma Miocene
Choerolophodon kisumuensis species	7	Animalia : Chordata : Mammalia : Proboscidea : Gomphotheriidae : Afrochoerodon : *Choerolophodon kisumuensis*	16.9 - 15.97 Ma Miocene
Gomphotherium angustidens species	7	Animalia : Chordata : Mammalia : Proboscidea : Gomphotheriidae : Gomphotherium : *Gomphotherium angustidens*	16.9 - 15.97 Ma Miocene
Tetrabelodon angustidens species	7	Animalia : Chordata : Mammalia : Proboscidea : Gomphotheriidae : Gomphotherium : *Tetrabelodon angustidens*	16.9 - 15.97 Ma Miocene
Mastodon spenceri species	7	Animalia : Chordata : Mammalia : Proboscidea : Mammutidae : Mammut : *Mastodon spenceri*	16.9 - 15.97 Ma Miocene
Zygolophodon aegyptensis species	7	Animalia : Chordata : Mammalia : Proboscidea : Mammutidae : Zygolophodon : *Zygolophodon aegyptensis*	16.9 - 15.97 Ma Miocene
Rimasuchus lloydi species	7	Animalia : Chordata : Reptilia : Crocodylia : Crocodylidae : Rimasuchus : *Rimasuchus lloydi*	16.9 - 15.97 Ma Miocene
Gavialis sp. genus	6	Animalia : Chordata : Reptilia : Crocodylia : Gavialidae : *Gavialis*	16.9 - 15.97 Ma Miocene
Tomistoma dowsoni species	7	Animalia : Chordata : Reptilia : Crocodylia : Crocodylidae : Tomistoma : *Tomistoma dowsoni*	16.9 - 15.97 Ma Miocene
Latentemys plowdeni species	7	Animalia : Chordata : Reptilia : Testudines : Podocnemididae : Latentemys : *Latentemys plowdeni*	16.9 - 15.97 Ma Miocene
Lemurchelys diasphax species	7	Animalia : Chordata : Reptilia : Testudines : Podocnemididae : Lemurchelys : *Lemurchelys diasphax*	23.03 - 15.97 Ma Miocene
Mogharemys blanckenhorni species	7	Animalia : Chordata : Reptilia : Testudines : Podocnemididae : Mogharemys : *Mogharemys blanckenhorni*	16.9 - 15.97 Ma Miocene
Podocnemis sp. genus	6	Animalia : Chordata : Reptilia : Testudines : Podocnemididae : *Podocnemis*	16.9 - 15.97 Ma Miocene
Trionychidae family	5	Animalia : Chordata : Reptilia : Testudines : Trionychidae	16.9 - 15.97 Ma Miocene

References

Stromer E. (1907) Fossile Wirbeltier-Reste aus dem Uadi Faregh und Uadi Natrun in Agypten, Amhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main 29, 97-132

Fourtau R. (1918) , Contribution à l'étude des vertébrés miocènes de l'Égypte, 1-109

Fortau R. (1920) Contribution a l'etude des vertebres Miocenes de l'Egypt, Egypt Survey Department, Cairo, 1-121

Miller E. R., Rasmussen D. T., et al (1997) Fossil storks (Ciconiidae) from the Late Eocene and Early Miocene of Egypt, The Ostrich 68 1, 23-26

Pickford M., Attia Y. S., et al (2001) Discovery of Prolibytherium magnieri Arambourg, 1961 (Artiodactyla, Climacoceratidae) in Egypt, Geodiversitas 23 4, 647-652

Sanders W. J., Miller E. R. (2002) New proboscideans from the early Miocene of Wadi Mogahara, Egypt, Journal of Vertebrate Paleontology 22 2, 388-404

Morlo M., Miller E. R., et al (2007) Creodonta and Carnivora from Wadi Moghra, Egypt, Journal of Vertebrate Paleontology 27 1, 145-159

Gaffney E. S., Meylan P. A., et al (2011) Evolution of the side-necked turtles: the family Podocnemididae, American Museum Novitates 350, 1-237

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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