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Ludlowville - Wanakah, New York, USA

Formation	Ludlowville
Member	Wanakah
Age:	388 - 383 Ma Devonian
Interval	Givetian
Lithology	mudstone, mudstone/"limestone", lime mudstone, "shale", "limestone"
Number of Collections	48
Number of Occurrences	608

Recorded Sample Locations

Location	Region	Stratigraphic Name
Locality 1	Erie County, New York, USA	Ludlowville - Wanakah
Locality 4	Erie County, New York, USA	Ludlowville - Wanakah
Locality 5	Erie County, New York, USA	Ludlowville - Wanakah
Locality 2	Erie County, New York, USA	Ludlowville - Wanakah
Locality 3	Erie County, New York, USA	Ludlowville - Wanakah
BCVS1-1	Erie County, New York, USA	Ludlowville - Wanakah
BCVS6-6	Erie County, New York, USA	Ludlowville - Wanakah
Locality 14	Genesee County, New York, USA	Ludlowville - Wanakah
Locality 19	Livingston County, New York, USA	Ludlowville - Wanakah

Associated Units

Stratigraphic Name	Age	Lithology	Occurrence Records
Hamilton - Ludlowville - Wanakah Shale	388 - 383 Ma Devonian	shale	11

Recorded Fossils

Accepted Name	Type	Hierarchy	Age
Tentaculites sp. genus	6	Animalia : Mollusca : Tentaculitidae : *Tentaculites*	387.7 - 382.7 Ma Devonian
Clionolithes sp. genus	6	*Clionolithes*	387.7 - 382.7 Ma Devonian
Spirorbis sp. genus	6	Animalia : Annelida : Polychaeta : Sabellida : Serpulidae : *Spirorbis*	387.7 - 382.7 Ma Devonian
Ostracoda class	3	Animalia : Arthropoda : Ostracoda	387.7 - 382.7 Ma Devonian
Euglyphella sp. genus	6	Animalia : Arthropoda : Ostracoda : Podocopida : Quasillitidae : *Euglyphella*	387.7 - 382.7 Ma Devonian
Ponderodictya sp. genus	6	Animalia : Arthropoda : Ostracoda : Podocopida : Quasillitidae : *Ponderodictya*	387.7 - 382.7 Ma Devonian
Greenops sp. genus	6	Animalia : Arthropoda : Trilobita : Phacopida : Acastidae : *Greenops*	387.7 - 382.7 Ma Devonian
Dipleura sp. genus	6	Animalia : Arthropoda : Trilobita : Phacopida : Bathycheilidae : *Dipleura*	387.7 - 382.7 Ma Devonian
Eldredgeops rana species	7	Animalia : Arthropoda : Trilobita : Phacopida : Phacopidae : Eldredgeops : *Eldredgeops rana*	387.7 - 382.7 Ma Devonian
Crania sp. genus	6	Animalia : Brachiopoda : Craniata : Craniida : Craniidae : *Crania*	387.7 - 382.7 Ma Devonian
Petrocrania sp. genus	6	Animalia : Brachiopoda : Craniata : Craniida : Craniidae : *Petrocrania*	387.7 - 382.7 Ma Devonian
Orbiculoidea sp. genus	6	Animalia : Brachiopoda : Lingulata : Lingulida : Discinidae : *Orbiculoidea*	387.7 - 382.7 Ma Devonian
Lingula sp. genus	6	Animalia : Brachiopoda : Lingulata : Lingulida : Lingulidae : *Lingula*	387.7 - 382.7 Ma Devonian
Pholidops sp. genus	6	Animalia : Brachiopoda : *Pholidops*	387.7 - 382.7 Ma Devonian
Athyris spiriferoides species	7	Animalia : Brachiopoda : Rhynchonellata : Athyridida : Athyrididae : Athyris : *Athyris spiriferoides*	387.7 - 382.7 Ma Devonian
Nucleospira concinna species	7	Animalia : Brachiopoda : Rhynchonellata : Athyridida : Nucleospiridae : Nucleospira : *Nucleospira concinna*	387.7 - 382.7 Ma Devonian
Atrypa reticularis species	7	Animalia : Brachiopoda : Rhynchonellata : Atrypida : Atrypidae : Atrypa : *Atrypa reticularis*	387.7 - 382.7 Ma Devonian
Tropidoleptus carinatus species	7	Animalia : Brachiopoda : Rhynchonellata : Terebratulida : Tropidoleptidae : Tropidoleptus : *Tropidoleptus carinatus*	387.7 - 382.7 Ma Devonian
Rhipidomella sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Orthida : Rhipidomellidae : *Rhipidomella*	387.7 - 382.7 Ma Devonian
Rhipidomella vanuxemi species	7	Animalia : Brachiopoda : Rhynchonellata : Orthida : Rhipidomellidae : Rhipidomella : *Rhipidomella vanuxemi*	387.7 - 382.7 Ma Devonian
Camarotoechia sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Rhynchonellida : Camarotoechiidae : *Camarotoechia*	387.7 - 382.7 Ma Devonian
Eumetabolotoechia sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Rhynchonellida : Leiorhynchidae : *Eumetabolotoechia*	387.7 - 382.7 Ma Devonian
Leiorhynchus sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Rhynchonellida : Leiorhynchidae : *Leiorhynchus*	387.7 - 382.7 Ma Devonian
Sapphicorhynchus sappho species	7	Animalia : Brachiopoda : Rhynchonellata : Rhynchonellida : Sapphicorhynchidae : Sapphicorhynchus : *Sapphicorhynchus sappho*	387.7 - 382.7 Ma Devonian
Ambocoelia umbonata species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Ambocoeliidae : Ambocoelia : *Ambocoelia umbonata*	387.7 - 382.7 Ma Devonian
Crurispina nana species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Ambocoeliidae : Crurispina : *Crurispina nana*	387.7 - 382.7 Ma Devonian
Elita sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Elythidae : *Elita*	387.7 - 382.7 Ma Devonian
Elita fimbriata species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Elythidae : Elita : *Elita fimbriata*	387.7 - 382.7 Ma Devonian
Mucrospirifer mucronatus species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Mucrospiriferidae : Mucrospirifer : *Mucrospirifer mucronatus*	387.7 - 382.7 Ma Devonian
Mediospirifer sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Spinocyrtiidae : *Mediospirifer*	387.7 - 382.7 Ma Devonian
Mediospirifer audaculus species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Spinocyrtiidae : Mediospirifer : *Mediospirifer audaculus*	387.7 - 382.7 Ma Devonian
Spinocyrtia granulosa species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferida : Spinocyrtiidae : Spinocyrtia : *Spinocyrtia granulosa*	387.7 - 382.7 Ma Devonian
Cyrtina sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Spiriferinida : Cyrtinidae : *Cyrtina*	387.7 - 382.7 Ma Devonian
Cyrtina hamiltonensis species	7	Animalia : Brachiopoda : Rhynchonellata : Spiriferinida : Cyrtinidae : Cyrtina : *Cyrtina hamiltonensis*	387.7 - 382.7 Ma Devonian
Centronella sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Terebratulida : Centronellidae : *Centronella*	387.7 - 382.7 Ma Devonian
Cryptonella sp. genus	6	Animalia : Brachiopoda : Rhynchonellata : Terebratulida : Cryptonellidae : *Cryptonella*	387.7 - 382.7 Ma Devonian
Schuchertella sp. genus	6	Animalia : Brachiopoda : Strophomenata : Orthotetida : Schuchertellidae : *Schuchertella*	387.7 - 382.7 Ma Devonian
Truncalosia sp. genus	6	Animalia : Brachiopoda : Strophomenata : Productida : Araksalosiidae : *Truncalosia*	387.7 - 382.7 Ma Devonian
Devonochonetes sp. genus	6	Animalia : Brachiopoda : Strophomenata : Productida : Chonetidae : *Devonochonetes*	387.7 - 382.7 Ma Devonian
Devonochonetes coronatus species	7	Animalia : Brachiopoda : Strophomenata : Productida : Chonetidae : Devonochonetes : *Devonochonetes coronatus*	387.7 - 382.7 Ma Devonian
Devonochonetes scitulus species	7	Animalia : Brachiopoda : Strophomenata : Productida : Chonetidae : Devonochonetes : *Devonochonetes scitulus*	387.7 - 382.7 Ma Devonian
Retichonetes (Sinochonetes) lepidus species	7	Animalia : Brachiopoda : Strophomenata : Productida : Chonetidae : Sinochonetes : *Retichonetes (Sinochonetes) lepidus*	387.7 - 382.7 Ma Devonian
Spinulicosta sp. genus	6	Animalia : Brachiopoda : Strophomenata : Productida : Productellidae : *Spinulicosta*	387.7 - 382.7 Ma Devonian
Douvillina sp. genus	6	Animalia : Brachiopoda : Strophomenata : Strophomenida : Douvillinidae : *Douvillina*	387.7 - 382.7 Ma Devonian
Protoleptostrophia perplana species	7	Animalia : Brachiopoda : Strophomenata : Strophomenida : Leptostrophiidae : Protoleptostrophia : *Protoleptostrophia perplana*	387.7 - 382.7 Ma Devonian
Megastrophia sp. genus	6	Animalia : Brachiopoda : Strophomenata : Strophomenida : Strophodontidae : *Megastrophia*	387.7 - 382.7 Ma Devonian
Pholidostrophia sp. genus	6	Animalia : Brachiopoda : Strophomenata : Strophomenida : Strophodontidae : *Pholidostrophia*	387.7 - 382.7 Ma Devonian
Bryozoa phylum	2	Animalia : Bryozoa	387.7 - 382.7 Ma Devonian
Hederella sp. genus	6	Animalia : Bryozoa : Stenolaemata : Reptariidae : *Hederella*	387.7 - 382.7 Ma Devonian
Acrogenia sp. genus	6	Animalia : Bryozoa : Stenolaemata : *Acrogenia*	387.7 - 382.7 Ma Devonian
Fenestrata order	4	Animalia : Bryozoa : Stenolaemata : Fenestrata	387.7 - 382.7 Ma Devonian
Paleschara sp. genus	6	Animalia : Bryozoa : Stenolaemata : *Paleschara*	387.7 - 382.7 Ma Devonian
Aulocystis sp. genus	6	Animalia : Cnidaria : Anthozoa : Auloporida : Aulocystidae : *Aulocystis*	387.7 - 382.7 Ma Devonian
Aulopora sp. genus	6	Animalia : Cnidaria : Anthozoa : Auloporidae : *Aulopora*	387.7 - 382.7 Ma Devonian
Cladochonus sp. genus	6	Animalia : Cnidaria : Anthozoa : Auloporidae : *Cladochonus*	387.7 - 382.7 Ma Devonian
Cystiphylloides sp. genus	6	Animalia : Cnidaria : Anthozoa : Digonophyllidae : *Cystiphylloides*	387.7 - 382.7 Ma Devonian
Favosites sp. genus	6	Animalia : Cnidaria : Anthozoa : Favositidae : *Favosites*	387.7 - 382.7 Ma Devonian
Michelinia stylopora species	7	Animalia : Cnidaria : Anthozoa : Favositida : Micheliniidae : Michelinia : *Michelinia stylopora*	387.7 - 382.7 Ma Devonian
Pleurodictyum sp. genus	6	Animalia : Cnidaria : Anthozoa : Micheliniidae : *Pleurodictyum*	387.7 - 382.7 Ma Devonian
Thamnoptychia sp. genus	6	Animalia : Cnidaria : Anthozoa : Pachyporidae : *Thamnoptychia*	387.7 - 382.7 Ma Devonian
Amplexiphyllum sp. genus	6	Animalia : Cnidaria : Anthozoa : Stereolasmatidae : *Amplexiphyllum*	387.7 - 382.7 Ma Devonian
Stereolasma rectum species	7	Animalia : Cnidaria : Anthozoa : Stauriida : Stereolasmatidae : Stereolasma : *Stereolasma rectum*	387.7 - 382.7 Ma Devonian
Heliophyllum halli species	7	Animalia : Cnidaria : Anthozoa : Zaphrentidae : Heliophyllum : *Heliophyllum halli*	387.7 - 382.7 Ma Devonian
Camerata subclass	3.3	Animalia : Echinodermata : Crinoidea : Camerata	387.7 - 382.7 Ma Devonian
Crinoidea class	3	Animalia : Echinodermata : Crinoidea	387.7 - 382.7 Ma Devonian
Deltacrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Deltacrinus*	387.7 - 382.7 Ma Devonian
Acanthocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : Rhodocrinitidae : *Acanthocrinus*	387.7 - 382.7 Ma Devonian
Cyttarocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Cyttarocrinus*	387.7 - 382.7 Ma Devonian
Aorocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : Coelocrinidae : *Aorocrinus*	387.7 - 382.7 Ma Devonian
Botryocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : Thalamocrinidae : *Botryocrinus*	387.7 - 382.7 Ma Devonian
Costalocrinus sentosus species	7	Animalia : Echinodermata : Crinoidea : Botryocrinidae : Costalocrinus : *Costalocrinus sentosus*	387.7 - 382.7 Ma Devonian
Anamesocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Anamesocrinus*	387.7 - 382.7 Ma Devonian
Ancyrocrinus bulbosus species	7	Animalia : Echinodermata : Crinoidea : Thalamocrinidae : Ancyrocrinus : *Ancyrocrinus bulbosus*	387.7 - 382.7 Ma Devonian
Atractocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Atractocrinus*	387.7 - 382.7 Ma Devonian
Bactrocrinites sp. genus	6	Animalia : Echinodermata : Crinoidea : *Bactrocrinites*	387.7 - 382.7 Ma Devonian
Kophinocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Kophinocrinus*	387.7 - 382.7 Ma Devonian
Zostocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Zostocrinus*	387.7 - 382.7 Ma Devonian
Decadocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : Decadocrinidae : *Decadocrinus*	387.7 - 382.7 Ma Devonian
Logocrinus geniculatus species	7	Animalia : Echinodermata : Crinoidea : Poteriocrinidae : Logocrinus : *Logocrinus geniculatus*	387.7 - 382.7 Ma Devonian
Gilbertsocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : Rhodocrinitidae : *Gilbertsocrinus*	387.7 - 382.7 Ma Devonian
Synaptocrinus sp. genus	6	Animalia : Echinodermata : Crinoidea : *Synaptocrinus*	387.7 - 382.7 Ma Devonian
Eutaxocrinus whiteavesi species	7	Animalia : Echinodermata : Crinoidea : Taxocrinidae : Eutaxocrinus : *Eutaxocrinus whiteavesi*	387.7 - 382.7 Ma Devonian
Heteroschisma sp. genus	6	Animalia : Echinodermata : Fissiculata : Codasteridae : *Heteroschisma*	387.7 - 382.7 Ma Devonian
Hyperoblastus sp. genus	6	Animalia : Echinodermata : Fissiculata : Phaenoschismatidae : *Hyperoblastus*	387.7 - 382.7 Ma Devonian
Placoblastus sp. genus	6	Animalia : Echinodermata : Spiraculata : *Placoblastus*	387.7 - 382.7 Ma Devonian
Dictyonema sp. genus	6	Animalia : Hemichordata : Pterobranchia : Dendrograptidae : *Dictyonema*	387.7 - 382.7 Ma Devonian
Modiella sp. genus	6	Animalia : Mollusca : Bivalvia : Cardiida : Kalenteridae : *Modiella*	387.7 - 382.7 Ma Devonian
Cypricardinia indenta species	7	Animalia : Mollusca : Bivalvia : Carditida : Cardiniidae : Cypricardinia : *Cypricardinia indenta*	387.7 - 382.7 Ma Devonian
Cypricardella sp. genus	6	Animalia : Mollusca : Bivalvia : Carditida : Astartidae : *Cypricardella*	387.7 - 382.7 Ma Devonian
Buchiola sp. genus	6	Animalia : Mollusca : Bivalvia : Praecardiidae : *Buchiola*	387.7 - 382.7 Ma Devonian
Modiomorpha sp. genus	6	Animalia : Mollusca : Bivalvia : Modiomorphida : Modiomorphidae : *Modiomorpha*	387.7 - 382.7 Ma Devonian
Pterochaenia sp. genus	6	Animalia : Mollusca : Bivalvia : Myalinida : Lunulacardiidae : *Pterochaenia*	387.7 - 382.7 Ma Devonian
Grammysia sp. genus	6	Animalia : Mollusca : Bivalvia : Grammysiidae : *Grammysia*	387.7 - 382.7 Ma Devonian
Pholadella sp. genus	6	Animalia : Mollusca : Bivalvia : *Pholadella*	387.7 - 382.7 Ma Devonian
Nuculites sp. genus	6	Animalia : Mollusca : Bivalvia : Nuculanida : Malletiidae : *Nuculites*	387.7 - 382.7 Ma Devonian
Palaeoneilo sp. genus	6	Animalia : Mollusca : Bivalvia : Nuculanida : Malletiidae : *Palaeoneilo*	387.7 - 382.7 Ma Devonian
Nucula sp. genus	6	Animalia : Mollusca : Bivalvia : Nuculanida : Nuculanidae : *Nucula*	387.7 - 382.7 Ma Devonian
Nuculoidea sp. genus	6	Animalia : Mollusca : Bivalvia : Nuculida : Nuculidae : *Nuculoidea*	387.7 - 382.7 Ma Devonian
Pterioidea superfamily	4.7	Animalia : Mollusca : Bivalvia : Ostreida : Pterioidea	387.7 - 382.7 Ma Devonian
Cornellites sp. genus	6	Animalia : Mollusca : Bivalvia : Ostreida : Pterineidae : *Cornellites*	387.7 - 382.7 Ma Devonian
Leptodesma (Leiopteria) sp. subgenus	6.3	Animalia : Mollusca : Bivalvia : Ostreida : Pterineidae : Leptodesma : *Leptodesma (Leiopteria)*	387.7 - 382.7 Ma Devonian
Ptychopteria sp. genus	6	Animalia : Mollusca : Bivalvia : Ostreida : Pterineidae : *Ptychopteria*	387.7 - 382.7 Ma Devonian
Orthonota sp. genus	6	Animalia : Mollusca : Bivalvia : Unionida : Iridinidae : *Orthonota*	387.7 - 382.7 Ma Devonian
Nautiloidea subclass	3.3	Animalia : Mollusca : Cephalopoda : Nautiloidea	387.7 - 382.7 Ma Devonian
Spyroceras sp. genus	6	Animalia : Mollusca : Cephalopoda : Pseudorthoceratidae : *Spyroceras*	387.7 - 382.7 Ma Devonian
Bellerophon leda species	7	Animalia : Mollusca : Gastropoda : Bellerophontidae : Retispira : *Bellerophon leda*	387.7 - 382.7 Ma Devonian
Platyceras sp. genus	6	Animalia : Mollusca : Gastropoda : Platyceratidae : *Platyceras*	387.7 - 382.7 Ma Devonian
Platyostoma lineata species	7	Animalia : Mollusca : Gastropoda : Euomphalina : Platyceratidae : Platyceras : *Platyostoma lineata*	387.7 - 382.7 Ma Devonian
Styliolina fissurella species	7	Animalia : Mollusca : Tentaculita : Styliolinida : Styliolinidae : Styliolina : *Styliolina fissurella*	387.7 - 382.7 Ma Devonian
Craniella hamiltoniae species	7	Animalia : Porifera : Demospongea : Astrophorida : Craniella : *Craniella hamiltoniae*	387.7 - 382.7 Ma Devonian

References

Clement C. R. (1982) The Occurrence and Paleoecology of Echinoderm Asseblages in the Ludlowville Shales (Middle Devonian) of western Erie County, New York, unpublished M.S. thesis, 1-110

Miller K. B. (1986) Depositional Environments and Sequences, "Pleurodictyum Zone," Ludlowville Formation of Western New York, Dynamic Stratigraphy and Depositional Environments of the Hamilton Group (Middle Devonian) in New York State, Part I, New York State Museum Bulletin 457, 57-77

Brett C. E., Dick V. B., et al (1991) Comparative Taphonomy and Paleoecology of Middle Devonian Dark Gray and Black Shale Facies from Western New York, Dynamic Stratigraphy and Depositional Environments of the Hamilton Group (Middle Devonian) in New York State, Part II; New York State Museum Bulletin 469, 5-36

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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